观赏植物花香性状形成及调控机制研究进展

doi:10.16420/j.issn.0513-353x.2025-0815

摘要/Abstract

摘要：

花香是植物、环境及生物体进行信息交流的重要信号，其主要由花部组织释放的多种挥发性有机化合物（VOC）构成，包括萜类、苯丙素/苯类化合物及脂肪酸衍生物等。花香物质的合成涉及多条代谢途径，且在不同发育阶段和组织部位呈现显著的时空动态特征，其生物合成受关键酶基因、转录因子、激素信号及环境因子的协同调控。以萜类化合物为主要香气成分的观赏植物中，萜类合成酶（TPS）家族和多个数量性状位点（QTL）决定了花香成分的多样性与特异性。随着多组学和分子育种技术的发展，转基因与基因编辑技术（如CRISPR/Cas9）在花香性状改良中展现出巨大潜力，同时，利用微生物“细胞工厂”合成天然产物为实现花香物质的规模化与可持续生产提供了新思路。本文中系统综述了植物花香的主要化学成分与代谢途径、释放的时空调节机制及其关键基因和调控网络，并展望了花香分子育种与产业化应用的发展方向。本综述旨在为花香形成的分子机制研究提供理论参考，并为其在观赏植物改良与相关产业中的应用提供新的视角。

关键词: 花香, 挥发性有机化合物, 代谢通路, 转录因子, 分子育种

Abstract:

Floral scent serves as a crucial signal mediating interactions between plants，their environment，and other organisms. It is predominantly composed of diverse volatile organic compounds（VOC）emitted from floral tissues，including terpenoids，phenylpropanoids/benzenoids，and fatty acid derivatives. The biosynthesis of floral volatiles involves multiple metabolic pathways and exhibits pronounced spatiotemporal dynamics across different developmental stages and floral tissues. This process is coordinately regulated by key enzyme-encoding genes，transcription factors，hormonal signaling，and environmental cues. In ornamental plants where terpenoids dominate the floral bouquet，members of the terpene synthase（TPS）gene family，together with several quantitative trait loci（QTL），largely determine the diversity and specificity of floral scent composition. With the rapid advancement of multi-omics and molecular breeding technologies，transgenic approaches and genome editing tools，such as CRISPR/Cas9，have shown great potential for modifying floral scent traits. Concurrently，microbial“cell factory” systems offer promising strategies for the scalable and sustainable production of natural floral volatiles. This review provides a comprehensive overview of the major chemical constituents and metabolic pathways underlying floral scent in plants，as well as the spatiotemporal regulation of volatile emission and the associated key genes and regulatory networks. Finally，it further discusses recent advances and future prospects in the molecular breeding and industrial applications of floral scent. The review aims to offer a theoretical framework for understanding the molecular mechanisms of scent formation and to provide new perspectives for its application in ornamental plant improvement and related industries.

Key words: floral scent, volatile organic compound, metabolic pathway, transcription factor, molecular breeding

王芳, 范燕萍. 观赏植物花香性状形成及调控机制研究进展[J]. 园艺学报, 2026, 53(2): 359-385.

WANG Fang, FAN Yanping. Molecular Insights into the Formation and Regulation of Floral Scent Traits in Ornamental Plants[J]. Acta Horticulturae Sinica, 2026, 53(2): 359-385.

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参考文献 180

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物种 Species	主要成分大类 Major classes of components	主要香气物质 Major aroma compounds	参考文献 Reference
金银花 Lonicera japonica	萜类 Terpenes	沉香醇、大牛儿烯 D Linalool，germacrene D	Feng et al.,2023
巴西番樱桃 Eugenia brasiliensis	芳香族、醇类、萜类 Aromatic compounds，alcohols，terpenes	苯甲醛、苯甲醇、2-苯乙醇、水杨酸甲酯、乙酸苄酯、1-辛醇、沉香醇 Benzaldehyde，benzyl alcohol，2-phenylethanol，methyl salicylate，benzyl acetate，1-octanol，linalool.	Cordeiro et al. 2019
鼓槌石斛 Dendrobium chrysotoxum	烯类、脂肪族酯 Alkenes，aliphatic esters	罗勒烯、乙酸辛酯 Ocimene，octyl acetate	赵瑞晶等,2024
建兰 Cymbidium ensifolium	脂肪酸类衍生物 Fatty acid derivatives	茉莉酸甲酯 Methyl jasmonate	高雪倩等,2025
文心兰 Oncidium Sharry Baby	萜类 Terpenes	罗勒烯、沉香醇 Ocimene，linalool	Yeh et al.,2022
杓兰 Cypripedium calceolus	萜类、脂肪族酯 Terpenes，aliphatic esters	沉香醇、乙酸辛酯 Linalool，octyl acetate	Braunschmid et al.,2017
蕙兰 Cymbidium‘Sael Bit’	脂肪族、芳香族 Aliphatic，aromatic	反式-2-辛烯醛、十三烷、苯 trans-2-Octenal，tridecane，benzene	Ramya et al.,2019
红掌 Anthurium andraeanum	萜类、芳香族酯 Terpenes，aromatic esters	桉油醇、乙酸苄酯 Eucalyptol，benzyl acetate	Wei et al.,2021
芍药 Paeonia lactiflora	萜类 Terpenes	香叶醇 Geraniol	Zhao et al.,2023
山茶 Camellia sinensis	萜类 Terpenes	沉香醇 Linalool	陈艺荃等,2024
昙花 Epiphyllum oxypetalum	萜类 Terpenes	香叶醇 Geraniol	Zhang et al.,2025
茉莉 Jasminum sambac	芳香族醛、芳香族酯、萜类 Aromatic aldehydes，aromatic esters，terpenes	苯乙醛、苯甲酸甲酯、乙酸苄酯、α-法尼烯 Benzaldehyde，methyl benzoate，benzyl acetate，α-farnesene	张宇航等,2025
紫丁香 Syringa vulgaris	萜类 Terpenes	罗勒烯 Ocimene	吴静等,2024
桂花 Osmanthus fragrans	萜类 Terpenes	β-紫罗兰酮、沉香醇 β-Ionone，linalool	Xin et al.,2013
百合 Lilium	萜类、芳香族酯 Terpenes，aromatic esters	1.8-桉油醇、沉香醇、β罗勒烯、苯甲酸甲酯 1,8-Cineole，linalool，β-ocimene，methyl benzoate	Kong et al.,2017
紫薇 Lagerstroemia indica	芳香族酚、萜类 Aromatic phenols，terpenes	丁香酚、香叶醇 Eugenol，geraniol	Zhou et al.,2024
姜花 Hedychium coronarium	萜类、芳香族酯 Terpenes，aromatic esters	（E）-β-罗勒烯、沉香醇和1,8-桉油醇、苯甲酸甲酯 (E)-β-Ocimene，linaloo，1,8-cineole，methyl benzoate	Yue et al.,2021；Zhou et al.,2021
铁线莲 Clematis florida	萜类 Terpenes	沉香醇 Linalool	Jiang et al.,2020
报春 Primula forbesii	萜类 Terpenes	沉香醇 Linalool	Jia et al.,2017
郁金香 Tulipa gesneriana	萜类、脂肪族类 Terpenes，aliphatic compounds	α-法呢烯、乙醇、十五烷、β-罗勒烯、昌叶烯、石竹烯、丙酮 α-Farnesene，ethanol，pentadecane，β-ocimene，longifolene，caryophyllene，acetone	Yang et al.,2020
玫瑰 Rosa rugosa	芳香族类、萜类、卤代烃、脂肪族类 Aromatic compounds，terpenes，halogenated hydrocarbons，aliphatic compounds	苯乙醇、香茅醇、二氯甲烷、正己烷、丙酮 2-Phenylethanol，citronellol，dichloromethane，n-hexane，acetone	Yang et al.,2020
梅花 Prunus mume	芳香族类 Aromatic compounds	乙酸苄酯、丁香酚 Benzyl acetate，eugenol	Zhang et al.,2019
黄兰 Magnolia champaca	萜类、脂肪族类 Terpenes，aliphatic compounds	β-沉香醇、2-甲基丁酸甲酯 β-Linalool，methyl 2-methylbutyrate	Dhandapani et al.,2017
蜡梅 Chimonanthus praecox	萜类 Terpenes	桃金娘烯醇 Myrtenol	Li et al.,2022
滇丁香 Luculia pinceana	芳香族类、萜类 Aromatic compounds ，terpenes	牡丹酚、α-法呢烯、（+）-环苜蓿烯、杜松烯 Paeonol，α-farnesene，cyclosativene，δ-cadinene	Li et al.,2016
巨花马兜铃 Aristolochia gigantea	萜类 Terpenes	柠檬醛、香茅醇、香茅醛 Citral，citronellol，citronellal	Martin et al.,2017
荷花 Nelumbo nucifera	芳香族类 Aromatic compounds	1,4-二甲氧基苯 1,4-Dimethoxybenzene	Wei et al.,2025a
晚香玉 Polianthes tuberosa	芳香族类、脂肪族类 Aromatic and aliphatic compounds	苯甲酸甲酯、苯甲酸苄酯、二十五烷、水杨酸甲酯 Methyl benzoate，benzyl benzoate，pentacosane，methyl salicylate	Kanani & Nazarideljou,2017
睡莲 Victoria cruziana	脂肪族类 Aliphatic compounds	己酸甲酯 Methyl hexanoate	Jiang et al.,2021
尾叶紫薇 Lagerstroemia caudata	萜类 Terpenes	橙花醇、香叶醇、沉香醇 Nerol，geraniol，linalool	Cai et al.,2023
紫藤 Wisteria sinenesis	萜类 Terpenes	（E）-β-罗勒烯、沉香醇 (E)-β-Ocimene，linalool	Jiang et al.,2011
金钱蒲 Acorus gramineus	呋喃类化合物 Furan compounds	2-（1E,3Z-己二烯基）四氢呋喃 2-(1E,3Z-hexadienyl)tetrahydrofuran	Azuma & Toyota,2012
澳洲香水石斛 Dendrobium ‘Aozhouxiangshui’	芳香族类、萜类 Aromatic compounds ，terpenes	1,4-二甲氧基苯、α-蒎烯、苯乙醇 1,4-Dimethoxybenzene，α-pinene，2-phenylethanol.	孔兰等,2024
野茉莉 Styrax japonicus	萜类 Terpenes	沉香醇、草蒿脑、大根香叶烯 Linalool，estragole，germacrene-d	Chen et al.,2021
大叶蝴蝶兰 Phalaenopsis bellina	萜类 Terpenes	沉香醇、香叶醇 Linalool，geraniol	Huang et al.,2021
七里香 Murraya paniculata	芳香族类、萜类 Aromatic compounds，terpenes	苯甲醛、苯乙醛、沉香醇、石竹烯、大根香叶烯、 α-法呢烯Benzaldehyde，phenylacetaldehyde，linaloo，caryophyllene，germacrene，α-farnesene	Datta et al.,2024
单花鸢尾 Iris uniflora	脂肪族类、芳香族类 Aliphatic and aromatic compounds	壬醛、癸醛、2,4-二叔丁基苯酚、十七烷 Nonyl aldehyde，capric aldehyde，2,4-di-tert-butylphenol，n-heptadecane	Cai et al.,2024
白花朱顶红 Hippeastrum brasilianum	萜类 Terpenes	（Z）-β-罗勒烯 (Z)-β-Ocimene	Meerow et al.,2017
绣球 Hydrangea arborescens	芳香族类 Aromatic compounds	苯乙酮 Acetophenone	Ke et al.,2023
夜来香 Cestrum nocturnum	芳香族类 Aromatic compounds	苯甲醛、苯乙醛、乙酸苄酯 Benzaldehyde，phenylacetaldehyde，benzyl acetate	Qiao et al.,2025
黄金国睡莲 Nymphaea ‘Eldorado’	芳香族类 Aromatic compounds	苯甲醛、苯甲醇、乙酸苄酯 Benzaldehyde，benzyl alcohol，benzyl acetate	Zhou et al.,2024
紫花含笑 Michelia crassipes	萜类 Terpenes	α-古芸烯、β-石竹烯、大根香叶烯B α-Copaene，β-Caryophyllene，Germacrene B	Yong et al.,2023
小叶丁香 Syringa pubescens	芳香族类	苯乙醛、苯乙醇 Phenylacetaldehyde，2-phenylethanol	Wei et al.,2025b
香豌豆 Lathyrus odoratus	萜类 Terpenes	α-香柠檬烯、沉香醇、α-荜澄茄油烯、香叶醇、 β-石竹烯、β-倍半水芹烯 α-Bergamotene，linalool，(−)-α-cubebene，geraniol，β-caryophyllene，β-sesquiphellandrene	Bao et al.,2020
德国鸢尾 Iris germanica	萜类 Terpenes	沉香醇 Linalool	Zhao et al.,2024

物种 Species	主要成分大类 Major classes of components	主要香气物质 Major aroma compounds	参考文献 Reference
金银花 Lonicera japonica	萜类 Terpenes	沉香醇、大牛儿烯 D Linalool，germacrene D	Feng et al.,2023
巴西番樱桃 Eugenia brasiliensis	芳香族、醇类、萜类 Aromatic compounds，alcohols，terpenes	苯甲醛、苯甲醇、2-苯乙醇、水杨酸甲酯、乙酸苄酯、1-辛醇、沉香醇 Benzaldehyde，benzyl alcohol，2-phenylethanol，methyl salicylate，benzyl acetate，1-octanol，linalool.	Cordeiro et al. 2019
鼓槌石斛 Dendrobium chrysotoxum	烯类、脂肪族酯 Alkenes，aliphatic esters	罗勒烯、乙酸辛酯 Ocimene，octyl acetate	赵瑞晶等,2024
建兰 Cymbidium ensifolium	脂肪酸类衍生物 Fatty acid derivatives	茉莉酸甲酯 Methyl jasmonate	高雪倩等,2025
文心兰 Oncidium Sharry Baby	萜类 Terpenes	罗勒烯、沉香醇 Ocimene，linalool	Yeh et al.,2022
杓兰 Cypripedium calceolus	萜类、脂肪族酯 Terpenes，aliphatic esters	沉香醇、乙酸辛酯 Linalool，octyl acetate	Braunschmid et al.,2017
蕙兰 Cymbidium‘Sael Bit’	脂肪族、芳香族 Aliphatic，aromatic	反式-2-辛烯醛、十三烷、苯 trans-2-Octenal，tridecane，benzene	Ramya et al.,2019
红掌 Anthurium andraeanum	萜类、芳香族酯 Terpenes，aromatic esters	桉油醇、乙酸苄酯 Eucalyptol，benzyl acetate	Wei et al.,2021
芍药 Paeonia lactiflora	萜类 Terpenes	香叶醇 Geraniol	Zhao et al.,2023
山茶 Camellia sinensis	萜类 Terpenes	沉香醇 Linalool	陈艺荃等,2024
昙花 Epiphyllum oxypetalum	萜类 Terpenes	香叶醇 Geraniol	Zhang et al.,2025
茉莉 Jasminum sambac	芳香族醛、芳香族酯、萜类 Aromatic aldehydes，aromatic esters，terpenes	苯乙醛、苯甲酸甲酯、乙酸苄酯、α-法尼烯 Benzaldehyde，methyl benzoate，benzyl acetate，α-farnesene	张宇航等,2025
紫丁香 Syringa vulgaris	萜类 Terpenes	罗勒烯 Ocimene	吴静等,2024
桂花 Osmanthus fragrans	萜类 Terpenes	β-紫罗兰酮、沉香醇 β-Ionone，linalool	Xin et al.,2013
百合 Lilium	萜类、芳香族酯 Terpenes，aromatic esters	1.8-桉油醇、沉香醇、β罗勒烯、苯甲酸甲酯 1,8-Cineole，linalool，β-ocimene，methyl benzoate	Kong et al.,2017
紫薇 Lagerstroemia indica	芳香族酚、萜类 Aromatic phenols，terpenes	丁香酚、香叶醇 Eugenol，geraniol	Zhou et al.,2024
姜花 Hedychium coronarium	萜类、芳香族酯 Terpenes，aromatic esters	（E）-β-罗勒烯、沉香醇和1,8-桉油醇、苯甲酸甲酯 (E)-β-Ocimene，linaloo，1,8-cineole，methyl benzoate	Yue et al.,2021；Zhou et al.,2021
铁线莲 Clematis florida	萜类 Terpenes	沉香醇 Linalool	Jiang et al.,2020
报春 Primula forbesii	萜类 Terpenes	沉香醇 Linalool	Jia et al.,2017
郁金香 Tulipa gesneriana	萜类、脂肪族类 Terpenes，aliphatic compounds	α-法呢烯、乙醇、十五烷、β-罗勒烯、昌叶烯、石竹烯、丙酮 α-Farnesene，ethanol，pentadecane，β-ocimene，longifolene，caryophyllene，acetone	Yang et al.,2020
玫瑰 Rosa rugosa	芳香族类、萜类、卤代烃、脂肪族类 Aromatic compounds，terpenes，halogenated hydrocarbons，aliphatic compounds	苯乙醇、香茅醇、二氯甲烷、正己烷、丙酮 2-Phenylethanol，citronellol，dichloromethane，n-hexane，acetone	Yang et al.,2020
梅花 Prunus mume	芳香族类 Aromatic compounds	乙酸苄酯、丁香酚 Benzyl acetate，eugenol	Zhang et al.,2019
黄兰 Magnolia champaca	萜类、脂肪族类 Terpenes，aliphatic compounds	β-沉香醇、2-甲基丁酸甲酯 β-Linalool，methyl 2-methylbutyrate	Dhandapani et al.,2017
蜡梅 Chimonanthus praecox	萜类 Terpenes	桃金娘烯醇 Myrtenol	Li et al.,2022
滇丁香 Luculia pinceana	芳香族类、萜类 Aromatic compounds ，terpenes	牡丹酚、α-法呢烯、（+）-环苜蓿烯、杜松烯 Paeonol，α-farnesene，cyclosativene，δ-cadinene	Li et al.,2016
巨花马兜铃 Aristolochia gigantea	萜类 Terpenes	柠檬醛、香茅醇、香茅醛 Citral，citronellol，citronellal	Martin et al.,2017
荷花 Nelumbo nucifera	芳香族类 Aromatic compounds	1,4-二甲氧基苯 1,4-Dimethoxybenzene	Wei et al.,2025a
晚香玉 Polianthes tuberosa	芳香族类、脂肪族类 Aromatic and aliphatic compounds	苯甲酸甲酯、苯甲酸苄酯、二十五烷、水杨酸甲酯 Methyl benzoate，benzyl benzoate，pentacosane，methyl salicylate	Kanani & Nazarideljou,2017
睡莲 Victoria cruziana	脂肪族类 Aliphatic compounds	己酸甲酯 Methyl hexanoate	Jiang et al.,2021
尾叶紫薇 Lagerstroemia caudata	萜类 Terpenes	橙花醇、香叶醇、沉香醇 Nerol，geraniol，linalool	Cai et al.,2023
紫藤 Wisteria sinenesis	萜类 Terpenes	（E）-β-罗勒烯、沉香醇 (E)-β-Ocimene，linalool	Jiang et al.,2011
金钱蒲 Acorus gramineus	呋喃类化合物 Furan compounds	2-（1E,3Z-己二烯基）四氢呋喃 2-(1E,3Z-hexadienyl)tetrahydrofuran	Azuma & Toyota,2012
澳洲香水石斛 Dendrobium ‘Aozhouxiangshui’	芳香族类、萜类 Aromatic compounds ，terpenes	1,4-二甲氧基苯、α-蒎烯、苯乙醇 1,4-Dimethoxybenzene，α-pinene，2-phenylethanol.	孔兰等,2024
野茉莉 Styrax japonicus	萜类 Terpenes	沉香醇、草蒿脑、大根香叶烯 Linalool，estragole，germacrene-d	Chen et al.,2021
大叶蝴蝶兰 Phalaenopsis bellina	萜类 Terpenes	沉香醇、香叶醇 Linalool，geraniol	Huang et al.,2021
七里香 Murraya paniculata	芳香族类、萜类 Aromatic compounds，terpenes	苯甲醛、苯乙醛、沉香醇、石竹烯、大根香叶烯、 α-法呢烯Benzaldehyde，phenylacetaldehyde，linaloo，caryophyllene，germacrene，α-farnesene	Datta et al.,2024
单花鸢尾 Iris uniflora	脂肪族类、芳香族类 Aliphatic and aromatic compounds	壬醛、癸醛、2,4-二叔丁基苯酚、十七烷 Nonyl aldehyde，capric aldehyde，2,4-di-tert-butylphenol，n-heptadecane	Cai et al.,2024
白花朱顶红 Hippeastrum brasilianum	萜类 Terpenes	（Z）-β-罗勒烯 (Z)-β-Ocimene	Meerow et al.,2017
绣球 Hydrangea arborescens	芳香族类 Aromatic compounds	苯乙酮 Acetophenone	Ke et al.,2023
夜来香 Cestrum nocturnum	芳香族类 Aromatic compounds	苯甲醛、苯乙醛、乙酸苄酯 Benzaldehyde，phenylacetaldehyde，benzyl acetate	Qiao et al.,2025
黄金国睡莲 Nymphaea ‘Eldorado’	芳香族类 Aromatic compounds	苯甲醛、苯甲醇、乙酸苄酯 Benzaldehyde，benzyl alcohol，benzyl acetate	Zhou et al.,2024
紫花含笑 Michelia crassipes	萜类 Terpenes	α-古芸烯、β-石竹烯、大根香叶烯B α-Copaene，β-Caryophyllene，Germacrene B	Yong et al.,2023
小叶丁香 Syringa pubescens	芳香族类	苯乙醛、苯乙醇 Phenylacetaldehyde，2-phenylethanol	Wei et al.,2025b
香豌豆 Lathyrus odoratus	萜类 Terpenes	α-香柠檬烯、沉香醇、α-荜澄茄油烯、香叶醇、 β-石竹烯、β-倍半水芹烯 α-Bergamotene，linalool，(−)-α-cubebene，geraniol，β-caryophyllene，β-sesquiphellandrene	Bao et al.,2020
德国鸢尾 Iris germanica	萜类 Terpenes	沉香醇 Linalool	Zhao et al.,2024

物种Species	基因Gene	合成产物Synthetic products	参考文献Reference
桂花 Osmanthus fragrans	OfTPS1，OfTPS2 OfTPS3，OfTPS7	参与合成沉香醇 Involved in linalool biosynthesis	Zeng et al.,2016
蕙兰 Cymbidium faberi	TPS18	催化GPP转化为β-月桂烯、香叶醇、α-蒎烯 Catalyzes the conversion of GPP into β-myrcene，geraniol，and α-pinene	Wang et al.,2021
铁线莲 Clematis florida	CfTPS1，CfTPS2	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Jiang et al.,2020
姜花 Hedychium coronarium	HcTPS1	催化GPP生成1,8-桉油醇和α-蒎烯 Catalyzes the conversion of GPP to 1,8-cineole and α-pinene	Yue et al.,2015,2021；Ke et al.,2021
	HcTPS5，HcTPS8	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
	HcTPS3	催化GPP生成（E）-β-罗勒烯 Catalyzes the conversion of GPP to (E)-β-ocimene
	HcTPS38	催化GPP生成香茅醇 Catalyzes the conversion of GPP to citronellol
	HcBSMT1	催化苯甲酸生成苯甲酸甲酯 Catalyzes the conversion of benzoic acid to methyl benzoate
	HcBSMT2	催化水杨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid to methyl salicylate
芍药 Paeonia lactiflora	PlTPS1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Zhao et al.,2025
	PlTPS4	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
	PlTPS6	催化FPP生成石竹烯和α-葎草烯 Catalyzes the conversion of FPP to caryophyllene and α-humulene
	PlTPS8	催化GPP生成α-蒎烯、沉香醇和2-蒈烯 and 2-pinene 催化FPP生成大根香叶烯、红没药烯 Catalyzes the conversion of FPP to germacrene and β-elemene
昙花 Epiphyllum oxypetalum	EoTPSa1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol 催化FPP生成法呢烯 Catalyzes the conversion of FPP to farnesene	Zhang et al.,2025
	EoTPSg3	催化GPP生成β-蒎烯、β罗勒烯、沉香醇等 etc 催化FPP生成古巴烯、法呢烯、金合欢醇等 Catalyzes the conversion of FPP to cubebene，farnesene，and nerolidol，etc
	EoTPSg1	催化GPP生成香叶醇、橙花醇、柠檬醛 Catalyzes the conversion of GPP to geraniol，nerol，and citral
紫丁香 Syringa oblata	SoTPS2，SoTPS3	参与合成罗勒烯 Involved in the biosynthesis of ocimene	吴静等,2024
西藏虎头兰 Cymbidium tracyanum	CtTPS2	催化GPP生成（E）-β-罗勒烯、α-松油醇、萜品油烯 Catalyzes the conversion of GPP to (E)-β-ocimene，α-terpineol，and terpinolene 催化FPP生成β-倍半水芹烯、α-姜黄烯、α-法呢烯、α-红没药烯、橙花叔醇 Catalyzes the conversion of FPP to β-germacrene，α-curcumene，α-farnesene，α-β-elemene，and nerolidol	Tu et al.,2025
	CtTPS3	催化GPP生成别罗勒烯 Catalyzes the conversion of GPP to allo-ocimene E）-金合欢醇 E)-nerolidol
	CtTPS8	催化GPP生成别罗勒烯和萜品油烯 Catalyzes the conversion of GPP to allo-ocimene and terpinolene 催化FPP生成（E）-β-石竹烯 Catalyzes the conversion of FPP to (E)-β-caryophyllene
黄精灵百合 Lilium‘Yelloween’	LiBSMT	催化苯甲酸生成苯甲酸甲酯 Catalyzes the conversion of benzoic acid to methyl benzoate 催化水杨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid to methyl salicylate	Wang et al.,2015
西伯利亚百合 Lilium‘Siberia’	LoTPS1	催化GPP生成（Z）-β-罗勒烯和（±）-沉香醇 Catalyzes the conversion of GPP to (Z)-β-ocimene and (±)-linalool	Abbas et al.,2019
	LoTPS3	催化GPP生成（±）-沉香醇 Catalyzes the conversion of GPP to（±）-linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Abbas et al.,2019
	LoAAT1	参与合成苯甲酸乙酯和苯甲酸甲酯 Involved in the biosynthesis of ethyl benzoate and methyl benzoate	Yue et al.,2023
	LoTPS2	催化FPP生成（E,E）-α-法呢烯 Catalyzes the conversion of FPP to (E,E)-α-farnesene	Abbas et al.,2020a
	LoTPS4	催化GPP生成D-柠檬烯、β-月桂烯 Catalyzes the conversion of GPP to D-limonene and β-myrcene	Abbas et al.,2020a
	LoTPS4	催化FPP生成反式-α-香柠檬烯 Catalyzes the conversion of FPP to trans-α-bergamotene
	LoTPS5	催化FPP生成角鲨烯 Catalyzes the conversion of FPP to squalene	Abbas et al.,2020b
	LiTPS2	催化GPP生成沉香醇、（E）-β-罗勒烯、月桂烯 Catalyzes the conversion of GPP to linalool，(E)-β-ocimene，and myrcene 催化FPP生成反式-橙花叔醇、（E）-β-法呢烯、α-法呢烯 Catalyzes the conversion of FPP to trans-nerolidol，(E)-β-farnesene，and α-farnesene	Zhang et al.,2020
西伯利亚百合 Lilium‘Siberia’	LiDXS	参与合成石竹烯 Involved in the biosynthesis of caryophyllene	Zhang et al.,2018
西伯利亚百合 Lilium‘Siberia’	LiDXR	参与合成沉香醇和石竹烯 Involved in the biosynthesis of linalool and caryophyllene	Zhang et al.,2018
黄栀 Gardenia jasminoides	GjTPS18	催化GPP生成α-松油醇、β-吡喃酮、D-柠檬烯、β-蒎烯 Catalyzes the conversion of GPP to α-terpineol，β-phellandrene，D-limonene，and β-pinene 催化FPP生成T-杜松醇、大香根叶烯D、古巴醇、α-卡地醇、勒多醇 Catalyzes the conversion of FPP to T-juniperol，germacrene D，cubebol，α-cadinol，and ledol	Chen et al.,2025
	GjTPS1	催化GPP生成β-罗勒烯 Catalyzes the conversion of GPP to β-ocimene 催化FPP生成α-法呢烯和橙花叔醇 Catalyzes the conversion of FPP to α-farnesene and nerolidol
	GjTPS27	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol
	GjTPS2，GjTPS3	催化GPP生成β-罗勒烯 Catalyzes the conversion of GPP to β-ocimene
	GjTPS19	催化FPP生成大香根叶烯D Catalyzes the conversion of FPP to germacrene D
大叶蝴蝶兰 Phalaenopsis bellina	PbTPS5，PbTPS9	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Huang et al.,2021
	PbTPS3	催化GPP生成沉香醇和罗勒烯 Catalyzes the conversion of GPP to linalool and ocimene
	PbTPS4	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
尾叶紫薇 Lagerstroemia caudata	LcTPS14	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Xu et al.,2024
姜荷花 Curcuma alismatifolia	CaTPS2	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Cao et al.,2024
姜荷花 Curcuma alismatifolia	CaTPS3	催化GPP生成β-月桂烯 Catalyzes the conversion of GPP to β-myrcene 催化FPP生成β-法呢烯 Catalyzes the conversion of FPP to β-farnesene	Cao et al.,2024
依兰 Cananga odorata	CoTPS1	催化GPP生成β-侧柏烯、桧烯、β-蒎烯、α-松油烯 Catalyzes the conversion of GPP to β-sabinene，thujene，β-pinene，α-terpinene	Jin et al.,2015
	CoTPS2	催化FPP生成β-金合欢烯、β-可巴烯、β-荜澄茄油烯 β-cubebenes
	CoTPS3	催化FPP生成α-香柠檬烯 Catalyzes the conversion of FPP to α-bergamotene
	CoTPS4	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol
素心腊梅 Chimonanthus praecox	CpTPS3	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool	Jiang et al.,2023
素心腊梅 Chimonanthus praecox	CpBEAT	以苯甲醇和乙酰辅酶A为底物合成乙酸苄酯 Synthesizes benzyl acetate from benzyl alcohol and acetyl-CoA as substrates	Jiang et al.,2023
香豌豆 Lathyrus odoratus	LoTPS4	催化GPP生成月桂烯、罗勒烯、异松油烯、沉香醇、香叶醇 Catalyzes the conversion of GPP to myrcene，ocimene，terpinolene，linalool，geraniol	Bao et al.,2020
香豌豆 Lathyrus odoratus	LoTPS7	催化GPP生成侧柏烯、β-蒎烯、β-水芹烯 β-germacrene	Bao et al.,2020
丹参 Salvia miltiorrhiza	SmTPS1	催化FPP生成β-榄香烯、（E）-β-石竹烯、α-葎草烯 Catalyzes the conversion of FPP to β-elemene，(E)-β-caryophyllene，α-humulene	Li et al.,2024
短舌匹菊 Pyrethrum parthenium	PpTPS5	催化FPP生成β-榄香烯 Catalyzes the conversion of FPP to β-elemene 催化GPP生成 β-月桂烯、D-柠檬烯、反式-β-罗勒烯、罗勒烯、蒈烯和沉香醇 Catalyzes the conversion of GPP to β-myrcene，D-limonene，trans-β-ocimene，ocimene，terpinene，and linalool	Wang et al.,2025b
德国鸢尾 Iris germanica	IgTPS14	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Zhao et al.,2024
茉莉花 Jasminum sambac	JsBEAT1 JsBEAT2	参与合成乙酸苄酯，异源表达增加矮牵牛中苯甲酸苄酯和乙酸苄酯含量 Involved in the biosynthesis of benzyl acetate，and its heterologous expression significantly increases the contents of benzyl benzoate and benzyl acetate in petunia	Wang et al.,2022
非洲茉莉 Stephanotis floribunda	SAMT	催化水杨酸和S-腺苷甲硫氨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid and S-adenosyl-L-methionine（SAM）into methyl salicylate	Pott et al.,2002
玫瑰 Rosa rugosa	RhAAT	参与合成苯乙酸苯乙酯和乙酸苄酯 Involved in the biosynthesis of phenethyl phenylacetate and benzyl acetate	Guterman et al.,2006
	RhNUDX1	参与合成香叶醇 Involved in the biosynthesis of geraniol	Magnard et al.,2015
	RhPAAS	参与合成2-苯乙醇 Involved in the biosynthesis of 2-phenylethanol	Roccia et al.,2019
	RcEGS1	病毒沉默RcEGS1降低丁香酚含量 Virus-induced silencing of RcEGS1 reduces eugenol content	Yan et al.,2018
	RrAAAT，RrPPDC1	异源转化增加矮牵牛中2-苯乙醇含量 Heterologous transformation increases the content of 2-phenylethanol in petunia	Sheng et al.,2018；Zeng et al.,2019
牡丹 Paeonia suffruticosa	PsuLIS	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool	Song,et al. 2024
晚香玉 Polianthes tuberosa	PtBCMT2	参与合成苯甲酸甲酯 Involved in the biosynthesis of methyl benzoate	Fan,et al. 2018
矮牵牛 Petunia hybrida	PhDEF	激活 EOBI 和 EOBII，调控苯丙烷类挥发性化合物的生物合成 Activation of EOBI and EOBII regulates the biosynthesis of phenylpropanoid volatile compounds	Bednarczyk et al.,2025
矮牵牛 Petunia hybrida	PhHSF19	激活PhPAL2启动子参与合成反式肉桂酸 Activation of the PhPAL2 promoter is involved in the biosynthesis of trans-cinnamic acid	Fu et al.,2022
小报春 Primula forbesii	PfDXS2	病毒沉默PfDXS2显著降低沉香醇的含量 Virus-induced silencing of PfDXS2 significantly reduces the content of linalool	Jia et al.,2023
克鲁兹王莲 Victoria cruziana	VcSABATH1， VcSABATH3	参与合成己酸甲酯 Involved in the biosynthesis of methyl hexanoate	Jiang et al.,2021
茶树 Camellia sinensis	CsAFS2	催化FPP生成α-法尼烯 Catalyzes the conversion of FPP to α-farnesene	Xu et al.,2025
月季‘月月粉’ Rosa chinensis‘Old Blush’	RcCCD4	裂解β-胡萝卜素参与合成二氢-β-紫罗兰酮 Cleavage of β-carotene is involved in the biosynthesis of dihydro-β-ionone	付琪等,2025
铁皮石斛 Dendrobium officinale	DoGES1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Zhao et al.,2020
梅花 Prunus mume	PmCAD2	催化肉桂醛生成肉桂醇 Catalyzes the conversion of cinnamaldehyde to cinnamyl alcohol	Zhang et al.,2022
水仙 Narcissus tazetta var. chinensis	NtHDR	烟草中过表达生成沉香醇、苯甲醇、苯乙醇 Overexpression in tobacco leads to the production of linalool，benzyl alcohol，and 2-phenylethanol	Hu et al.,2023

物种Species	基因Gene	合成产物Synthetic products	参考文献Reference
桂花 Osmanthus fragrans	OfTPS1，OfTPS2 OfTPS3，OfTPS7	参与合成沉香醇 Involved in linalool biosynthesis	Zeng et al.,2016
蕙兰 Cymbidium faberi	TPS18	催化GPP转化为β-月桂烯、香叶醇、α-蒎烯 Catalyzes the conversion of GPP into β-myrcene，geraniol，and α-pinene	Wang et al.,2021
铁线莲 Clematis florida	CfTPS1，CfTPS2	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Jiang et al.,2020
姜花 Hedychium coronarium	HcTPS1	催化GPP生成1,8-桉油醇和α-蒎烯 Catalyzes the conversion of GPP to 1,8-cineole and α-pinene	Yue et al.,2015,2021；Ke et al.,2021
	HcTPS5，HcTPS8	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
	HcTPS3	催化GPP生成（E）-β-罗勒烯 Catalyzes the conversion of GPP to (E)-β-ocimene
	HcTPS38	催化GPP生成香茅醇 Catalyzes the conversion of GPP to citronellol
	HcBSMT1	催化苯甲酸生成苯甲酸甲酯 Catalyzes the conversion of benzoic acid to methyl benzoate
	HcBSMT2	催化水杨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid to methyl salicylate
芍药 Paeonia lactiflora	PlTPS1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Zhao et al.,2025
	PlTPS4	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
	PlTPS6	催化FPP生成石竹烯和α-葎草烯 Catalyzes the conversion of FPP to caryophyllene and α-humulene
	PlTPS8	催化GPP生成α-蒎烯、沉香醇和2-蒈烯 and 2-pinene 催化FPP生成大根香叶烯、红没药烯 Catalyzes the conversion of FPP to germacrene and β-elemene
昙花 Epiphyllum oxypetalum	EoTPSa1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol 催化FPP生成法呢烯 Catalyzes the conversion of FPP to farnesene	Zhang et al.,2025
	EoTPSg3	催化GPP生成β-蒎烯、β罗勒烯、沉香醇等 etc 催化FPP生成古巴烯、法呢烯、金合欢醇等 Catalyzes the conversion of FPP to cubebene，farnesene，and nerolidol，etc
	EoTPSg1	催化GPP生成香叶醇、橙花醇、柠檬醛 Catalyzes the conversion of GPP to geraniol，nerol，and citral
紫丁香 Syringa oblata	SoTPS2，SoTPS3	参与合成罗勒烯 Involved in the biosynthesis of ocimene	吴静等,2024
西藏虎头兰 Cymbidium tracyanum	CtTPS2	催化GPP生成（E）-β-罗勒烯、α-松油醇、萜品油烯 Catalyzes the conversion of GPP to (E)-β-ocimene，α-terpineol，and terpinolene 催化FPP生成β-倍半水芹烯、α-姜黄烯、α-法呢烯、α-红没药烯、橙花叔醇 Catalyzes the conversion of FPP to β-germacrene，α-curcumene，α-farnesene，α-β-elemene，and nerolidol	Tu et al.,2025
	CtTPS3	催化GPP生成别罗勒烯 Catalyzes the conversion of GPP to allo-ocimene E）-金合欢醇 E)-nerolidol
	CtTPS8	催化GPP生成别罗勒烯和萜品油烯 Catalyzes the conversion of GPP to allo-ocimene and terpinolene 催化FPP生成（E）-β-石竹烯 Catalyzes the conversion of FPP to (E)-β-caryophyllene
黄精灵百合 Lilium‘Yelloween’	LiBSMT	催化苯甲酸生成苯甲酸甲酯 Catalyzes the conversion of benzoic acid to methyl benzoate 催化水杨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid to methyl salicylate	Wang et al.,2015
西伯利亚百合 Lilium‘Siberia’	LoTPS1	催化GPP生成（Z）-β-罗勒烯和（±）-沉香醇 Catalyzes the conversion of GPP to (Z)-β-ocimene and (±)-linalool	Abbas et al.,2019
	LoTPS3	催化GPP生成（±）-沉香醇 Catalyzes the conversion of GPP to（±）-linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Abbas et al.,2019
	LoAAT1	参与合成苯甲酸乙酯和苯甲酸甲酯 Involved in the biosynthesis of ethyl benzoate and methyl benzoate	Yue et al.,2023
	LoTPS2	催化FPP生成（E,E）-α-法呢烯 Catalyzes the conversion of FPP to (E,E)-α-farnesene	Abbas et al.,2020a
	LoTPS4	催化GPP生成D-柠檬烯、β-月桂烯 Catalyzes the conversion of GPP to D-limonene and β-myrcene	Abbas et al.,2020a
	LoTPS4	催化FPP生成反式-α-香柠檬烯 Catalyzes the conversion of FPP to trans-α-bergamotene
	LoTPS5	催化FPP生成角鲨烯 Catalyzes the conversion of FPP to squalene	Abbas et al.,2020b
	LiTPS2	催化GPP生成沉香醇、（E）-β-罗勒烯、月桂烯 Catalyzes the conversion of GPP to linalool，(E)-β-ocimene，and myrcene 催化FPP生成反式-橙花叔醇、（E）-β-法呢烯、α-法呢烯 Catalyzes the conversion of FPP to trans-nerolidol，(E)-β-farnesene，and α-farnesene	Zhang et al.,2020
西伯利亚百合 Lilium‘Siberia’	LiDXS	参与合成石竹烯 Involved in the biosynthesis of caryophyllene	Zhang et al.,2018
西伯利亚百合 Lilium‘Siberia’	LiDXR	参与合成沉香醇和石竹烯 Involved in the biosynthesis of linalool and caryophyllene	Zhang et al.,2018
黄栀 Gardenia jasminoides	GjTPS18	催化GPP生成α-松油醇、β-吡喃酮、D-柠檬烯、β-蒎烯 Catalyzes the conversion of GPP to α-terpineol，β-phellandrene，D-limonene，and β-pinene 催化FPP生成T-杜松醇、大香根叶烯D、古巴醇、α-卡地醇、勒多醇 Catalyzes the conversion of FPP to T-juniperol，germacrene D，cubebol，α-cadinol，and ledol	Chen et al.,2025
	GjTPS1	催化GPP生成β-罗勒烯 Catalyzes the conversion of GPP to β-ocimene 催化FPP生成α-法呢烯和橙花叔醇 Catalyzes the conversion of FPP to α-farnesene and nerolidol
	GjTPS27	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol
	GjTPS2，GjTPS3	催化GPP生成β-罗勒烯 Catalyzes the conversion of GPP to β-ocimene
	GjTPS19	催化FPP生成大香根叶烯D Catalyzes the conversion of FPP to germacrene D
大叶蝴蝶兰 Phalaenopsis bellina	PbTPS5，PbTPS9	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Huang et al.,2021
	PbTPS3	催化GPP生成沉香醇和罗勒烯 Catalyzes the conversion of GPP to linalool and ocimene
	PbTPS4	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool
尾叶紫薇 Lagerstroemia caudata	LcTPS14	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Xu et al.,2024
姜荷花 Curcuma alismatifolia	CaTPS2	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Cao et al.,2024
姜荷花 Curcuma alismatifolia	CaTPS3	催化GPP生成β-月桂烯 Catalyzes the conversion of GPP to β-myrcene 催化FPP生成β-法呢烯 Catalyzes the conversion of FPP to β-farnesene	Cao et al.,2024
依兰 Cananga odorata	CoTPS1	催化GPP生成β-侧柏烯、桧烯、β-蒎烯、α-松油烯 Catalyzes the conversion of GPP to β-sabinene，thujene，β-pinene，α-terpinene	Jin et al.,2015
	CoTPS2	催化FPP生成β-金合欢烯、β-可巴烯、β-荜澄茄油烯 β-cubebenes
	CoTPS3	催化FPP生成α-香柠檬烯 Catalyzes the conversion of FPP to α-bergamotene
	CoTPS4	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol
素心腊梅 Chimonanthus praecox	CpTPS3	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool	Jiang et al.,2023
素心腊梅 Chimonanthus praecox	CpBEAT	以苯甲醇和乙酰辅酶A为底物合成乙酸苄酯 Synthesizes benzyl acetate from benzyl alcohol and acetyl-CoA as substrates	Jiang et al.,2023
香豌豆 Lathyrus odoratus	LoTPS4	催化GPP生成月桂烯、罗勒烯、异松油烯、沉香醇、香叶醇 Catalyzes the conversion of GPP to myrcene，ocimene，terpinolene，linalool，geraniol	Bao et al.,2020
香豌豆 Lathyrus odoratus	LoTPS7	催化GPP生成侧柏烯、β-蒎烯、β-水芹烯 β-germacrene	Bao et al.,2020
丹参 Salvia miltiorrhiza	SmTPS1	催化FPP生成β-榄香烯、（E）-β-石竹烯、α-葎草烯 Catalyzes the conversion of FPP to β-elemene，(E)-β-caryophyllene，α-humulene	Li et al.,2024
短舌匹菊 Pyrethrum parthenium	PpTPS5	催化FPP生成β-榄香烯 Catalyzes the conversion of FPP to β-elemene 催化GPP生成 β-月桂烯、D-柠檬烯、反式-β-罗勒烯、罗勒烯、蒈烯和沉香醇 Catalyzes the conversion of GPP to β-myrcene，D-limonene，trans-β-ocimene，ocimene，terpinene，and linalool	Wang et al.,2025b
德国鸢尾 Iris germanica	IgTPS14	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool 催化FPP生成橙花叔醇 Catalyzes the conversion of FPP to nerolidol	Zhao et al.,2024
茉莉花 Jasminum sambac	JsBEAT1 JsBEAT2	参与合成乙酸苄酯，异源表达增加矮牵牛中苯甲酸苄酯和乙酸苄酯含量 Involved in the biosynthesis of benzyl acetate，and its heterologous expression significantly increases the contents of benzyl benzoate and benzyl acetate in petunia	Wang et al.,2022
非洲茉莉 Stephanotis floribunda	SAMT	催化水杨酸和S-腺苷甲硫氨酸生成水杨酸甲酯 Catalyzes the conversion of salicylic acid and S-adenosyl-L-methionine（SAM）into methyl salicylate	Pott et al.,2002
玫瑰 Rosa rugosa	RhAAT	参与合成苯乙酸苯乙酯和乙酸苄酯 Involved in the biosynthesis of phenethyl phenylacetate and benzyl acetate	Guterman et al.,2006
	RhNUDX1	参与合成香叶醇 Involved in the biosynthesis of geraniol	Magnard et al.,2015
	RhPAAS	参与合成2-苯乙醇 Involved in the biosynthesis of 2-phenylethanol	Roccia et al.,2019
	RcEGS1	病毒沉默RcEGS1降低丁香酚含量 Virus-induced silencing of RcEGS1 reduces eugenol content	Yan et al.,2018
	RrAAAT，RrPPDC1	异源转化增加矮牵牛中2-苯乙醇含量 Heterologous transformation increases the content of 2-phenylethanol in petunia	Sheng et al.,2018；Zeng et al.,2019
牡丹 Paeonia suffruticosa	PsuLIS	催化GPP生成沉香醇 Catalyzes the conversion of GPP to linalool	Song,et al. 2024
晚香玉 Polianthes tuberosa	PtBCMT2	参与合成苯甲酸甲酯 Involved in the biosynthesis of methyl benzoate	Fan,et al. 2018
矮牵牛 Petunia hybrida	PhDEF	激活 EOBI 和 EOBII，调控苯丙烷类挥发性化合物的生物合成 Activation of EOBI and EOBII regulates the biosynthesis of phenylpropanoid volatile compounds	Bednarczyk et al.,2025
矮牵牛 Petunia hybrida	PhHSF19	激活PhPAL2启动子参与合成反式肉桂酸 Activation of the PhPAL2 promoter is involved in the biosynthesis of trans-cinnamic acid	Fu et al.,2022
小报春 Primula forbesii	PfDXS2	病毒沉默PfDXS2显著降低沉香醇的含量 Virus-induced silencing of PfDXS2 significantly reduces the content of linalool	Jia et al.,2023
克鲁兹王莲 Victoria cruziana	VcSABATH1， VcSABATH3	参与合成己酸甲酯 Involved in the biosynthesis of methyl hexanoate	Jiang et al.,2021
茶树 Camellia sinensis	CsAFS2	催化FPP生成α-法尼烯 Catalyzes the conversion of FPP to α-farnesene	Xu et al.,2025
月季‘月月粉’ Rosa chinensis‘Old Blush’	RcCCD4	裂解β-胡萝卜素参与合成二氢-β-紫罗兰酮 Cleavage of β-carotene is involved in the biosynthesis of dihydro-β-ionone	付琪等,2025
铁皮石斛 Dendrobium officinale	DoGES1	催化GPP生成香叶醇 Catalyzes the conversion of GPP to geraniol	Zhao et al.,2020
梅花 Prunus mume	PmCAD2	催化肉桂醛生成肉桂醇 Catalyzes the conversion of cinnamaldehyde to cinnamyl alcohol	Zhang et al.,2022
水仙 Narcissus tazetta var. chinensis	NtHDR	烟草中过表达生成沉香醇、苯甲醇、苯乙醇 Overexpression in tobacco leads to the production of linalool，benzyl alcohol，and 2-phenylethanol	Hu et al.,2023

物种 Species	转录因子家族 Transcription factor	基因 Gene	作用机制 Regulatory mechanism	参考文献 Reference
矮牵牛 Petunia hybrida	R2R3-MYB	ODO1 EBOI EBOII	控制苯丙素类/苯类化合物的合成 Regulates the biosynthesis of phenylpropanoid/phenolic compounds	Spitzer-Rimon et al.,2012
	ERF	PhERF6	沉默PhERF6显著增加苯类/苯丙烷类物质的含量 Silencing PhERF6 significantly increases the content of phenolic/phenylpropanoid compounds	Liu et al.,2017
	bHLH	PhbHLH19	激活PhPAL2启动子参与合成苯类/苯丙烷 Activation of the PhPAL2 promoter is involved in the biosynthesis of phenolic/phenylpropanoid compounds	Niu et al.,2017
芍药 Paeonia lactiflora	R2R3-MYB	PlMYB73 PlMYB108 PlMYB70	激活PlTPS1的表达参与合成香叶醇 Activation of PlTPS1 expression is involved in the biosynthesis of geraniol	Zhao et al.,2023
棉花 Gossypium arboreum	WRKY	GaWRKY1	结合倍半萜环化酶CAD1-A启动子催化合成棉酚 Binding to the CAD1-A promoter of sesquiterpene cyclase catalyzes the biosynthesis of gossypol	Xu et al.,2004
文心兰Oncidium	MYB	CCA1	控制花香释放Regulates floral scent emission	Yeh et al.,2022
蝴蝶兰 Phalaenopsis orchids	bHLH	PbbHLH4	参与单萜生物合成 Involved in monoterpene biosynthesis	Chuang et al.,2018
姜花 Hedychium coronarium	MYB	HcMYB1	与HcIAA4蛋白相互作用调控花香产生 Interacts with HcIAA4 protein to regulate floral scent production	Ke et al.,2021；Abbas et al.,2021a
	MYB	HcMYB2	激活HcBSMT2表达参与合成苯甲酸甲酯 Activation of HcTPS5 expression is involved in the biosynthesis of linalool；Activation of HcBSMT2 expression is involved in the biosynthesis of methyl benzoate	Ke et al.,2021；Abbas et al.,2021a
	ARF	HcARF5	与HcIAA4、HcIAA6和HcMYB1互作参与β-罗勒烯合成 Interacts with HcIAA4，HcIAA6，and HcMYB1 to participate in the biosynthesis of β-ocimene	Abbas et al.,2021b
蕙兰 Cymbidium ‘Sael Bit’	MYB	CsMYB1	参与合成聚丙烯酸酯和2-甲基丁醛 Involved in the biosynthesis of polyacrylate and 2-methylbutanal	Ramya et al.,2019
茶树 Camellia sinensis	NAC	CsNAC29	激活CsAFS2启动子参与合成α-法呢烯 Activation of the CsAFS2 promoter is involved in the biosynthesis of α-farnesene	Xu et al.,2025
桂花 Osmanthus fragrans	bHLH	OfbHLH35	激活OfTPS2启动子参与合成沉香醇 Activation of the OfTPS2 promoter is involved in the biosynthesis of linalool	Han et al.,2022
	AP2/ERF	OfERF2	激活OfCCD1和OfCCD4启动子参与合成紫罗兰酮 Activation of the OfCCD1 and OfCCD4 promoters is involved in the biosynthesis of ionone	Han et al.,2022
	YABBY	OfYABBY12	负调控OfCCD4表达，促进合成β-紫罗兰酮 Negative regulation of OfCCD4 expression promotes the biosynthesis of β-ionone	Shi et al.,2024
	R2R3-MYB	OfMYB21	激活OfTPS2启动子参与合成沉香醇 Activation of the OfTPS2 promoter is involved in the biosynthesis of linalool	Lan et al.,2023
薰衣草 Lavandula angustifolia	bHLH	LaMYC4	烟草中异源表达可增加石竹烯含量 Heterologous expression in tobacco increases caryophyllene content	Dong et al.,2022
温郁金 Curcuma wenyujin	bHLH	CwMYC2	瞬时表达会增加β-榄香烯的合成 Transient expression enhances the biosynthesis of β-elemene	Liu et al.,2023
蜡梅 Chimonanthus praecox	R2R3-MYB	CpODO1	激活CpCYP71启动子参与合成苯甲醇 Activation of the CpCYP71 promoter is involved in the biosynthesis of benzyl alcohol	Yong et al.,2025
西伯利亚百合 Lilium‘Siberia’	NAC	LiNAC100	激活LiLiS启动子参与合成沉香醇 Activation of the LiLiS promoter is involved in the biosynthesis of linalool	Liu et al.,2024
西伯利亚百合 Lilium‘Siberia’	MYB	LiMYB305	病毒沉默降低沉香醇、罗勒烯和月桂烯含量 Virus-induced silencing reduces the contents of linalool，ocimene，and myrcene	Yang et al.,2022
金鱼草 Antirrhinum majus	MYB	AmMYB24	激活AmOCS启动子参与合成罗勒烯 Activation of the AmOCS promoter is involved in the biosynthesis of ocimene	Han et al.,2022
玫瑰 Rosa rugosa	bHLH	RrbHLH105	激活RrTPS5和RrTPS8启动子负调控香叶醇含量 Activation of RrTPS5 and RrTPS8 promoters negatively regulates geraniol content	Bao et al.,2025
除虫菊 Tanacetum cinerariifolium	bHLH	TcMYC2	参与合成除虫菊酯 Involved in the biosynthesis of pyrethrins	Zeng et al.,2022
短舌匹菊 Pyrethrum parthenium	MYB	PpMYB9	激活PpTPS5启动子参与萜类物质合成 Activation of the PpTPS5 promoter is involved in terpenoid biosynthesis	Wang et al.,2025