园艺学报 ›› 2025, Vol. 52 ›› Issue (1): 73-87.doi: 10.16420/j.issn.0513-353x.2023-0632
李芮, 王稳, 杜明辉, 刘根忠, 马方放*(), 包志龙*(
)
收稿日期:
2024-08-29
修回日期:
2024-10-25
出版日期:
2025-01-25
发布日期:
2025-01-18
通讯作者:
基金资助:
LI Rui, WANG Wen, DU Minghui, LIU Genzhong, MA Fangfang*(), BAO Zhilong*(
)
Received:
2024-08-29
Revised:
2024-10-25
Published:
2025-01-25
Online:
2025-01-18
Contact:
* E-mail:zbao@sdau.edu.cn,fma@sdau.edu.cn
摘要:
BONZAI1(BON1)是高度保守的钙依赖膜结合蛋白Copine家族成员,具有调控生长发育的重要作用。番茄SlBON1与拟南芥BON1的蛋白序列相似度为66.84%。通过CRISPR-Cas9技术获得两个番茄SlBON1敲除突变体cr6和cr8,分别为61个碱基缺失突变和1个碱基插入突变。与野生型相比,SlBON1敲除突变体的地上和地下部生长均受到显著抑制,下胚轴中核内周期下降,叶片光合速率显著下降,淀粉含量升高。转录组分析显示SlBON1敲除突变体中激素信号转导信号通路、MAPK信号通路以及细胞周期的差异基因富集度较高。推测SlBON1通过调控植物激素代谢、细胞周期及光合作用等促进了番茄植株的营养生长。
李芮, 王稳, 杜明辉, 刘根忠, 马方放, 包志龙. SlBON1调控番茄植株营养生长机制的研究[J]. 园艺学报, 2025, 52(1): 73-87.
LI Rui, WANG Wen, DU Minghui, LIU Genzhong, MA Fangfang, BAO Zhilong. Mechanistic Studies on SlBON1-Regulated Plant Vegetative Growth in Tomato[J]. Acta Horticulturae Sinica, 2025, 52(1): 73-87.
用途 Usage | 引物名称 Primer name | 引物序列(5′-3′) Primer sequence |
---|---|---|
CRISPR/Cas9载体构建 Construct of vector | Solyc03g068840-BsFL-F | ATATATGGTCTCGATTGGAATGTCCACCGTTCTCGTGTTTTAGAGCTAGAAATAGC |
Solyc03g068840-BsRL-R | ATTATTGGTCTCGAAACCGATTACTTGGCATGGGGCCAATCTCTTAGTCGACTCTAC | |
靶点验证引物 Target validation primer | BON1-2-T1-F | TTTGGACTGACGCAGCTAGT |
BON1-2-T2-R | TCTGGTGACACCTACACCTG | |
载体验证引物 Vector validation primer | U6-26p-F | TGTCCCAGGATTAGAATGATTAGGC |
U6-29p-R | AGCCCTCTTCTTTCGATCCATCAAC | |
实时荧光定量PCR qRT-PCR | SlBON1-2-qPCR-F | TCGCGATGTGCTTTCCAAGA |
SlBON1-2-qPCR-R | CCACTTGGGATTCAATGAGTTTGG | |
内参基因 Internal reference gene | SlCBL1-qRT-F SlCBL1-qRT-R | CCATCCAAATGCTCCGATCGATGA TGCCTCTCAATGAAGCCTTGTTGC |
表1 本研究中所用的引物序列
Table 1 Primer sequences used in this study
用途 Usage | 引物名称 Primer name | 引物序列(5′-3′) Primer sequence |
---|---|---|
CRISPR/Cas9载体构建 Construct of vector | Solyc03g068840-BsFL-F | ATATATGGTCTCGATTGGAATGTCCACCGTTCTCGTGTTTTAGAGCTAGAAATAGC |
Solyc03g068840-BsRL-R | ATTATTGGTCTCGAAACCGATTACTTGGCATGGGGCCAATCTCTTAGTCGACTCTAC | |
靶点验证引物 Target validation primer | BON1-2-T1-F | TTTGGACTGACGCAGCTAGT |
BON1-2-T2-R | TCTGGTGACACCTACACCTG | |
载体验证引物 Vector validation primer | U6-26p-F | TGTCCCAGGATTAGAATGATTAGGC |
U6-29p-R | AGCCCTCTTCTTTCGATCCATCAAC | |
实时荧光定量PCR qRT-PCR | SlBON1-2-qPCR-F | TCGCGATGTGCTTTCCAAGA |
SlBON1-2-qPCR-R | CCACTTGGGATTCAATGAGTTTGG | |
内参基因 Internal reference gene | SlCBL1-qRT-F SlCBL1-qRT-R | CCATCCAAATGCTCCGATCGATGA TGCCTCTCAATGAAGCCTTGTTGC |
图1 SlBON1的系统进化树(A)、基因表达(B)、蛋白结构(C)、基因结构(D)和敲除突变类型(E) 采用Duncan’s法进行差异显著性分析。不同小写字母表示差异显著,P < 0.05。下同
Fig. 1 SlBON1 phylogenetic tree(A),tissue-specific expression(B),protein structure(C),gene structure(D),and knockout mutations(E) The different lowercase letters above the column indicate the statistical significance determined by Duncan’s multiple range test(P < 0.05)
图2 番茄SlBON1敲除株系(cr6、cr8)与野生型(WT)的下胚轴表型、长度、直径和核内周期 EI:核内周期指数;2C ~ 32C:细胞倍性
Fig. 2 The morphology,length,diameter and endoreduplication of hypocotyls in the wild type and SlBON1 knockout mutants cr6 and cr8 EI:Endoreduplication index. 2C-32C:Cell ploidy
图3 番茄70 d苗龄SlBON1敲除株系(cr6、cr8)与野生型(WT)的植株表型、地上部鲜、干质量、株高、茎粗和叶片面积
Fig. 3 Plant morphology,fresh and dry weight of aboveground tissues,plant height,stem diameter and leaf area in 70-day old seedlings of the wild type and SlBON1 knockout mutants cr6 and cr8
图4 番茄70 d苗龄SlBON1敲除株系(cr6、cr8)与野生型(WT)的光合指标、叶绿素和淀粉含量
Fig. 4 The photosynthetic parameters,chlorophyll and starch content in 70 day old seedlings of the wild type and SlBON1 knockout mutants cr6 and cr8
图5 番茄70d苗龄SlBON1敲除株系(cr6、cr8)与野生型(WT)植株超氧化物歧化酶、过氧化氢酶和过氧化物酶活性
Fig. 5 The enzyme activities of superoxide dismutase,catalase and peroxidase in 70-day old seedlings of the wild type and SlBON1 knockout mutants cr6 and cr8
图6 番茄70 d苗龄SlBON1敲除株系(cr6、cr8)与野生型(WT)的根系表型、鲜、干质量、根长、表面积和分支数量
Fig. 6 The morphology,fresh and dry weight,length,surface area and branch number of roots in 70-day old seedlings of the wild type and SlBON1 knockout mutants cr6 and cr8
图7 番茄野生型(WT)与SlBON1敲除株系cr6和cr8差异基因热图
Fig. 7 The heatmap analysis of differentially expressed genes in the wild type and SlBON1 knockout mutants cr6 and cr8
图8 番茄野生型(WT)与SlBON1敲除株系(cr6和cr8)差异表达基因KEGG通路分析
Fig. 8 KEGG pathway analysis of differentially expressed genes in the wild type and SlBON1 knockout mutants cr6 and cr8
图9 番茄70 d苗龄SlBON1敲除株系(cr6、cr8)与野生型(WT)4条信号通路差异表达基因热图分析
Fig. 9 The heatmap analysis of differentially expressed genes of four pathways in 70-day old seedlings of the wild type and SlBON1 knockout mutants cr6 and cr8
基因类别 Gene category | 基因ID Gene ID | 功能注释 Function |
---|---|---|
丝裂原活化蛋白激酶信号通路(上调)MAPK signaling pathway(up) | Solyc03g078360.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶LECRK1 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK1 |
Solyc07g053130.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g03230亚型X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g03230 isoform X1 | |
Solyc09g075910.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶 LECRK4 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK4 | |
Solyc07g055670.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶SD2-5 G-type lectin S-receptor-like serine/threonine-protein kinase SD2-5 | |
Solyc05g008310.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 | |
Solyc07g063770.3.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 isoform X1 | |
Solyc10g006710.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 isoform X1 | |
Solyc11g005630.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶LECRK3,部分序列 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK3,partial | |
Solyc04g077270.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 | |
Solyc07g053120.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g03230亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g03230 isoform X1 | |
Solyc12g006840.2.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶CES101 G-type lectin S-receptor-like serine/threonine-protein kinase CES101 | |
Solyc08g077560.4.1 | 可能的丝氨酸/苏氨酸蛋白激酶PBL19 Probable serine/threonine-protein kinase PBL19 | |
Solyc12g009310.2.1 | 可能的丝氨酸/苏氨酸蛋白激酶PBL17 Probable serine/threonine-protein kinase PBL17 | |
Solyc04g082500.4.1 | 可能的丝氨酸/苏氨酸蛋白激酶PIX13 亚型 X2 Probable serine/threonine-protein kinase PIX13 isoform X2 | |
Solyc11g020230.1.1 | 丝氨酸/苏氨酸蛋白激酶样蛋白CCR4 Serine/threonine-protein kinase-like protein CCR4 | |
Solyc09g008010.3.1 | 丝氨酸/苏氨酸蛋白激酶PCRK1亚型X2 Serine/threonine-protein kinase PCRK1 isoform X2 | |
Solyc01g007990.4.1 | 富含半胱氨酸的受体样蛋白激酶2亚型X2 Cysteine-rich receptor-like protein kinase 2 isoform X2 | |
Solyc06g071810.1.1 | 富亮氨酸重复类受体激酶 Leucine-rich repeat receptor-like kinase | |
Solyc07g006480.4.1 | 可能是无活性的富含亮氨酸重复序列的类受体蛋白激酶At5g48380 Probably inactive leucine-rich repeat receptor-like protein kinase At5g48380 | |
Solyc12g096710.2.1 | 可能是无活性的富含亮氨酸重复序列的类受体蛋白激酶At5g48380 Probably inactive leucine-rich repeat receptor-like protein kinase At5g48380 | |
Solyc02g080080.3.1 | 假定受体样蛋白激酶At4g00960 亚型 X1Putative receptor-like protein kinase At4g00960 isoform X1 | |
Solyc02g080010.2.1 | 假定受体样蛋白激酶At4g00960 亚型 X1Putative receptor-like protein kinase At4g00960 isoform X1 | |
Solyc02g080070.3.1 | 假定受体样蛋白激酶At4g00960 Putative receptor-like protein kinase At4g00960 | |
Solyc01g109950.4.1 | 假定受体样蛋白激酶At4g39110 Probable receptor-like protein kinase At4g39110 | |
Solyc01g067020.4.1 | 非典型受体样激酶1前体 Atypical receptor-like kinase 1 precursor | |
Solyc09g011330.3.1 | 假定受体蛋白激酶ZmPK1 Putative receptor protein kinase ZmPK1 | |
Solyc02g089900.1.1 | LysM结构域类受体激酶4 LysM domain receptor-like kinase 4 | |
Solyc09g010850.4.1 | 受体样胞质激酶176亚型X2 Receptor-like cytoplasmic kinase 176 isoform X2 | |
Solyc10g084250.1.1 | CladeⅧ凝集素受体激酶 Clade VIII lectin receptor kinase | |
Solyc07g007020.3.1 | 壁相关受体激酶样1 Wall-associated receptor kinase-like 1 | |
Solyc02g090110.3.1 | 壁相关受体激酶样14前体 Wall-associated receptor kinase-like 14 precursor | |
Solyc02g079600.3.1 | 抗锈病激酶Lr10亚型X1 Rust resistance kinase Lr10 isoform X1 | |
Solyc11g007430.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc04g007380.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc11g007410.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc05g008960.3.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc04g007390.4.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc05g008980.2.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc05g008950.4.1 | 叶锈病10抗病位点类受体蛋白激酶1.1亚型x1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 isoform x1 | |
油菜素甾醇 BR | Solyc02g085360.4.1(up) | 细胞色素P450 90B3 Cytochrome P450 90B3 |
Solyc10g086500.1.1 (down) | 类固醇5α还原酶DET2 Steroid 5 alpha reductase DET2 | |
细胞周期(上 调)Cell cycle (up) | Solyc01g079500.4.1 | DNA复制许可因子MCM7样蛋白 DNA replication licensing factor MCM7-like protein |
Solyc07g005020.3.1 | DNA复制许可因子MCM5 DNA replication licensing factor MCM5 | |
Solyc01g110130.3.1 | DNA复制许可因子MCM4 DNA replication licensing factor MCM4 | |
Solyc06g073510.3.1 | 复制复合体亚基2的起点 Origin of replication complex subunit 2 | |
Solyc06g076860.4.1 | 细胞分裂控制蛋白6同系物B Cell division control protein 6 homolog B | |
Solyc12g088520.2.1 | 假定细胞周期蛋白A3-3 Putative cyclin-A3-3 | |
Solyc12g088530.2.1 | 假定细胞周期蛋白A3-1 Putative cyclin-A3-1 | |
植物激素信号(下调) Plant hormone signal transduction (down) | Solyc01g110590.4.1 | 生长素响应蛋白SAUR50-like Auxin-responsive protein SAUR50-like |
Solyc01g110880.1.1 | 生长素响应蛋白SAUR21-like Auxin-responsive protein SAUR21-like | |
Solyc11g011660.1.1 | 生长素响应蛋白SAUR68-like Auxin-responsive protein SAUR68-like | |
Solyc06g008030.3.1 | 转录因子PIF1亚型X1 Transcription factor PIF1 isoform X1 | |
Solyc01g102300.3.1 | 转录因子PIF3亚型X1 Transcription factor PIF3 isoform X1 | |
Solyc01g106040.3.1 | GATA转录因子19样异构体2 GATA transcription factor 19-like isoform X2 | |
Solyc10g076920.3.1 | 脱落酸不敏感5样蛋白2 Abscisic acid-insensitive 5-like protein 2 | |
Solyc09g089610.3.1 | 乙烯受体 Ethylene receptor | |
Solyc03g007230.4.1 | 蛋白磷酸酶2C 51-like Protein phosphatase 2C 51-like | |
Solyc03g096670.3.1 | 蛋白磷酸酶2C AHG3同源物 Protein phosphatase 2C AHG3 homolog | |
Solyc05g052980.4.1 | 蛋白磷酸酶2C 37 Protein phosphatase 2C 37 |
表2 4条信号通路差异表达基因列表
Table 2 List of differentially expressed genes in four pathways
基因类别 Gene category | 基因ID Gene ID | 功能注释 Function |
---|---|---|
丝裂原活化蛋白激酶信号通路(上调)MAPK signaling pathway(up) | Solyc03g078360.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶LECRK1 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK1 |
Solyc07g053130.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g03230亚型X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g03230 isoform X1 | |
Solyc09g075910.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶 LECRK4 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK4 | |
Solyc07g055670.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶SD2-5 G-type lectin S-receptor-like serine/threonine-protein kinase SD2-5 | |
Solyc05g008310.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 | |
Solyc07g063770.3.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 isoform X1 | |
Solyc10g006710.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 isoform X1 | |
Solyc11g005630.1.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶LECRK3,部分序列 G-type lectin S-receptor-like serine/threonine-protein kinase LECRK3,partial | |
Solyc04g077270.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g27290 G-type lectin S-receptor-like serine/threonine-protein kinase At4g27290 | |
Solyc07g053120.4.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶At4g03230亚型 X1 G-type lectin S-receptor-like serine/threonine-protein kinase At4g03230 isoform X1 | |
Solyc12g006840.2.1 | G型凝集素S受体样丝氨酸/苏氨酸蛋白激酶CES101 G-type lectin S-receptor-like serine/threonine-protein kinase CES101 | |
Solyc08g077560.4.1 | 可能的丝氨酸/苏氨酸蛋白激酶PBL19 Probable serine/threonine-protein kinase PBL19 | |
Solyc12g009310.2.1 | 可能的丝氨酸/苏氨酸蛋白激酶PBL17 Probable serine/threonine-protein kinase PBL17 | |
Solyc04g082500.4.1 | 可能的丝氨酸/苏氨酸蛋白激酶PIX13 亚型 X2 Probable serine/threonine-protein kinase PIX13 isoform X2 | |
Solyc11g020230.1.1 | 丝氨酸/苏氨酸蛋白激酶样蛋白CCR4 Serine/threonine-protein kinase-like protein CCR4 | |
Solyc09g008010.3.1 | 丝氨酸/苏氨酸蛋白激酶PCRK1亚型X2 Serine/threonine-protein kinase PCRK1 isoform X2 | |
Solyc01g007990.4.1 | 富含半胱氨酸的受体样蛋白激酶2亚型X2 Cysteine-rich receptor-like protein kinase 2 isoform X2 | |
Solyc06g071810.1.1 | 富亮氨酸重复类受体激酶 Leucine-rich repeat receptor-like kinase | |
Solyc07g006480.4.1 | 可能是无活性的富含亮氨酸重复序列的类受体蛋白激酶At5g48380 Probably inactive leucine-rich repeat receptor-like protein kinase At5g48380 | |
Solyc12g096710.2.1 | 可能是无活性的富含亮氨酸重复序列的类受体蛋白激酶At5g48380 Probably inactive leucine-rich repeat receptor-like protein kinase At5g48380 | |
Solyc02g080080.3.1 | 假定受体样蛋白激酶At4g00960 亚型 X1Putative receptor-like protein kinase At4g00960 isoform X1 | |
Solyc02g080010.2.1 | 假定受体样蛋白激酶At4g00960 亚型 X1Putative receptor-like protein kinase At4g00960 isoform X1 | |
Solyc02g080070.3.1 | 假定受体样蛋白激酶At4g00960 Putative receptor-like protein kinase At4g00960 | |
Solyc01g109950.4.1 | 假定受体样蛋白激酶At4g39110 Probable receptor-like protein kinase At4g39110 | |
Solyc01g067020.4.1 | 非典型受体样激酶1前体 Atypical receptor-like kinase 1 precursor | |
Solyc09g011330.3.1 | 假定受体蛋白激酶ZmPK1 Putative receptor protein kinase ZmPK1 | |
Solyc02g089900.1.1 | LysM结构域类受体激酶4 LysM domain receptor-like kinase 4 | |
Solyc09g010850.4.1 | 受体样胞质激酶176亚型X2 Receptor-like cytoplasmic kinase 176 isoform X2 | |
Solyc10g084250.1.1 | CladeⅧ凝集素受体激酶 Clade VIII lectin receptor kinase | |
Solyc07g007020.3.1 | 壁相关受体激酶样1 Wall-associated receptor kinase-like 1 | |
Solyc02g090110.3.1 | 壁相关受体激酶样14前体 Wall-associated receptor kinase-like 14 precursor | |
Solyc02g079600.3.1 | 抗锈病激酶Lr10亚型X1 Rust resistance kinase Lr10 isoform X1 | |
Solyc11g007430.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc04g007380.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc11g007410.3.1 | 抗锈病激酶Lr10 Rust resistance kinase Lr10 | |
Solyc05g008960.3.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc04g007390.4.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc05g008980.2.1 | 叶锈病10抗病位点类受体蛋白激酶1.1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 | |
Solyc05g008950.4.1 | 叶锈病10抗病位点类受体蛋白激酶1.1亚型x1 Leaf rust 10 disease-resistance locus receptor-like protein kinase-like 1.1 isoform x1 | |
油菜素甾醇 BR | Solyc02g085360.4.1(up) | 细胞色素P450 90B3 Cytochrome P450 90B3 |
Solyc10g086500.1.1 (down) | 类固醇5α还原酶DET2 Steroid 5 alpha reductase DET2 | |
细胞周期(上 调)Cell cycle (up) | Solyc01g079500.4.1 | DNA复制许可因子MCM7样蛋白 DNA replication licensing factor MCM7-like protein |
Solyc07g005020.3.1 | DNA复制许可因子MCM5 DNA replication licensing factor MCM5 | |
Solyc01g110130.3.1 | DNA复制许可因子MCM4 DNA replication licensing factor MCM4 | |
Solyc06g073510.3.1 | 复制复合体亚基2的起点 Origin of replication complex subunit 2 | |
Solyc06g076860.4.1 | 细胞分裂控制蛋白6同系物B Cell division control protein 6 homolog B | |
Solyc12g088520.2.1 | 假定细胞周期蛋白A3-3 Putative cyclin-A3-3 | |
Solyc12g088530.2.1 | 假定细胞周期蛋白A3-1 Putative cyclin-A3-1 | |
植物激素信号(下调) Plant hormone signal transduction (down) | Solyc01g110590.4.1 | 生长素响应蛋白SAUR50-like Auxin-responsive protein SAUR50-like |
Solyc01g110880.1.1 | 生长素响应蛋白SAUR21-like Auxin-responsive protein SAUR21-like | |
Solyc11g011660.1.1 | 生长素响应蛋白SAUR68-like Auxin-responsive protein SAUR68-like | |
Solyc06g008030.3.1 | 转录因子PIF1亚型X1 Transcription factor PIF1 isoform X1 | |
Solyc01g102300.3.1 | 转录因子PIF3亚型X1 Transcription factor PIF3 isoform X1 | |
Solyc01g106040.3.1 | GATA转录因子19样异构体2 GATA transcription factor 19-like isoform X2 | |
Solyc10g076920.3.1 | 脱落酸不敏感5样蛋白2 Abscisic acid-insensitive 5-like protein 2 | |
Solyc09g089610.3.1 | 乙烯受体 Ethylene receptor | |
Solyc03g007230.4.1 | 蛋白磷酸酶2C 51-like Protein phosphatase 2C 51-like | |
Solyc03g096670.3.1 | 蛋白磷酸酶2C AHG3同源物 Protein phosphatase 2C AHG3 homolog | |
Solyc05g052980.4.1 | 蛋白磷酸酶2C 37 Protein phosphatase 2C 37 |
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