国兰育种：历史、现状和展望

doi:10.16420/j.issn.0513-353x.2024-0946

摘要/Abstract

摘要：

国兰是中国传统文化的物质载体，栽培历史悠久，市场前景广阔。20世纪80年代以前，国兰育种采用引种驯化和选择育种方法进行。近年来，国兰在有性杂交育种、诱变育种、多倍体育种、细胞工程育种和分子育种等方面取得了重要进展，培育出一批花大、观花期长、易繁殖、适合产业化、大众化和国际化发展需求的新品种。本文综述了国兰育种的历史和研究进展，展望了育种未来，为推动国兰育种和产业高质量发展提供参考。

关键词: 国兰, 育种, 杂交, 组培快繁

Abstract:

Chinese Cymbidium，a symbol of traditional Chinese culture，has a long cultivation history and promising market prospects. Before the 1980s，breeding of Chinese Cymbidium was primarily achieved through introduction and domestication and selection breeding. In recent years，significant progress has been made in areas such as sexual hybridization breeding，mutation breeding，polyploid breeding，cell engineering breeding and molecular breeding. As a result，dozens of new cultivars have been developed，characterized by large flowers，long florescence，high micropropagation efficiency and suitability for industrialization，popularization，and international development. This article reviews the history of Chinese Cymbidium breeding，summarizes key advancements，and explores future prospects，offering valuable insights for promoting the high-quality development of both breeding and its associated industry.

Key words: Chinese Cymbidium, breeding, hybridization, micropropagation

贺子怡, 曾瑞珍, 谢利, 魏倩, 张志胜. 国兰育种：历史、现状和展望[J]. 园艺学报, 2025, 52(8): 2233-2248.

HE Ziyi, ZENG Ruizhen, XIE Li, WEI Qian, and ZHANG Zhisheng. Breeding of Chinese Cymbidium：History，Current Status and Future Prospects[J]. Acta Horticulturae Sinica, 2025, 52(8): 2233-2248.

图/表 4

参考文献 54

[55]	Park H Y, Kang K W, Kim D H, Sivanesan I. 2018. In vitro propagation of Cymbidium goeringii Reichenbach fil. through direct adventitious shoot regeneration. Physiology and Molecular Biology of Plants, 24 (2):307-313.
[56]	Pan Hong. 2008. Preliminary research on techniques of radiation induced mutation and tissue culture techniques in Cymbidium[M. D. Dissertation]. Fuzhou: Fujian Agriculture and Forestry University. (in Chinese)
	潘宏. 2008. 兰花辐射诱变与组织培养技术初步研究[硕士论文]. 福州: 福建农林大学.
[57]	Peng M X, Chen R M, Wei Q, Guo H R, Zeng R Z, Xie L, Zhang Z S, Chen J J. 2023. Effects of genotype,light,and plant growth regulators on rhizome browning,proliferation,and sprouting in Cymbidium. HortScience, 58 (6):671-676.
[58]	Shin Y H, Song I J, Kang E J, Bae T H, Sun H J, Kang S Y, Lim P O, Lee H Y. 2011. Development of leaf mutant cultivars of Cymbidium goeringii by Ethylmethanesulfonate(EMS)treatment. Korean Journal of Plant Resources, 24 (1):17-22.
[59]	Song Dandan. 2021. Effect of leaf color and pattern mutation on micropropagation and growth characteristics of Cymbidium‘Xiao Feng’[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	宋丹丹. 2021. 叶艺突变对‘小凤兰’组培快繁和生长特性的影响[硕士论文]. 广州: 华南农业大学.
[60]	Su Yan. 2016. Studies on rapid propagation of hybrid offspring of Cymbidium kanran and changes of physiology and biochemistry[M. D. Dissertation]. Fuzhou: Fujian Agriculture and Forestry University. (in Chinese)
	苏妍. 2016. 寒兰杂交后代快速繁殖体系及生理生化研究[硕士论文]. 福州：福建农林大学.
[61]	Sun Chongbo, Xiang Lin, Li Bojun, Qin Dehui, Li Xiaobai. 2012. A new Cymbidium goeringii cultivar‘Fuwamei’. Acta Horticulturae Sinica, 39 (12):2549-2551. (in Chinese)
	孙崇波, 向林, 李伯钧, 秦德辉, 李小白. 2012. 春兰新品种‘福娃梅’. 园艺学报, 39 (12):2549-2551.
[62]	Sun Ye, Bao Jianzhong, Zhang Tian, Ma Hui, Liu Chungui, Li Fengtong, Yuan Yuan, Chen Xiulan. 2018. A new Cymbidium goeringii cultivar‘Yanghongmei’. Acta Horticulturae Sinica, 45 (S2):2801-2802. (in Chinese)
	孙叶, 包建忠, 张甜, 马辉, 刘春贵, 李风童, 袁媛, 陈秀兰. 2018. 春兰新品种‘扬红梅’. 园艺学报, 45 (S2):2801-2802.
[1]	Ahmad S, Chen J L, Chen G Z, Huang J, Hao Y, Shi X L, Liu Y Y, Tu S, Zhou Y Z, Zhao K, Lan S R, Liu Z J, Peng D H. 2022. Transcriptional proposition for uniquely developed protocorm flowering in three orchid species:Resources for innovative breeding. Frontiers in Plant Sciences,13:942591.
[2]	Balilashaki K, Vahedi M, Ho T T, Niu S C, Cardoso J C, Zotz G, Khodamzadeh A A. 2022. Biochemical,cellular and molecular aspects of Cymbidium orchids:An ecological and economic overview. Acta Physiologiae Plantarum, 44 (2):1-17.
[3]	Bao Jianzhong, Sun Ye, Li Fengtong, Liu Chungui, Yuan Yuan, Ma Hui, Zhang Tian, Chen Xiulan. 2017. Breeding and cultivation techniques of new Cymbidium goeringii cultivars‘Yellow butterfly’and‘Ruihe’. Chinese Horticulturae Abstracts, 33 (10):1-3,16. (in Chinese)
	包建忠, 孙叶, 李风童, 刘春贵, 袁媛, 马辉, 张甜, 陈秀兰. 2017. 春兰新品种‘黄蝴蝶’‘瑞荷’的选育与栽培技术. 中国园艺文摘, 33 (10):1-3,16.
[4]	Bao Jianzhong, Sun Ye, Liu Chungui, Ma Hui, Chen Xiulan. 2010. Preliminary report on the breeding of new germplasm by irradiation mutagenesis of orchid. Jiangsu Agricultural Sciences,(6):269-270. (in Chinese)
	包建忠, 孙叶, 刘春贵, 马辉, 陈秀兰. 2010. 兰辐照诱变新种质选育研究初报. 江苏农业科学,(6):269-270.
[5]	Bu Zhaoyang, He Jingzhou, Huang Changyan, Zeng Yanhua. 2019. A new Cymbidium cultivar‘Sweet Princess’. Acta Horticulturae Sinica, 46 (S2):2875-2876. (in Chinese)
	卜朝阳, 何荆洲, 黄昌艳, 曾艳华. 2019. 兰花新品种‘香公主’. 园艺学报, 46 (S2):2875-2876.
[6]	Chen Hua, Lin Bing, Pan Hong. 2005. Preliminary report on the mutagenesis effect of Chinese Cymbidium. Fujian Agricultural Science and Technology,(4):24-25. (in Chinese)
	陈华, 林兵, 潘宏. 2005. 国兰辐射诱变效应研究初报. 福建农业科技,(4):24-25.
[7]	Chen Xiaomei. 2008. Studies on rapid propagation and tetraploid breeding of Cymbidium faberi Rolfe[M. D. Dissertation]. Chongqing: Southwest University. (in Chinese)
	陈晓梅. 2008. 蕙兰离体快繁及四倍体培育研究[硕士论文]. 重庆: 西南大学.
[63]	Sun Ye, Liu Chungui, Li Fengtong, Ma Hui, Zhang Jing, Chen Xiulan, Bao Jianzhong. 2016. Mutagenic effects of ⁶⁰Coγ irradiation on Cymbidium goeringii‘Xueshan’. Molecular Plant Breeding, 14 (10):2762-2768. (in Chinese)
	孙叶, 刘春贵, 李风童, 马辉, 张靓, 陈秀兰, 包建忠. 2016. ⁶⁰Coγ射线辐照对春兰‘雪山’的诱变效应. 分子植物育种, 14 (10):2762-2768.
[64]	Vilcherrez-Atoche J A, Iiyama C M, Cardoso J C. 2022. Polyploidization in orchids:From cellular changes to breeding applications. Plants, 11 (4):469.
[65]	Wang Buqiong, Li Zhilin, Yu Chaoxiu. 2005. Progress on orchid breeding. Acta Horticulturae Sinica, 32 (3):551-556. (in Chinese)
	王卜琼, 李枝林, 余朝秀. 2005. 兰花育种研究进展. 园艺学报, 32 (3):551-556.
[66]	Wang Jian, Le Chaoyin, Xie Wei, Liang Wei, Lin Zhi, Guo Zhenghong. 2006. RAPD molecular markers of Cymbidium fragrance related genes. Jiangsu Agricultural Sciences,(5):78-79. (in Chinese)
	王健, 乐超银, 谢伟, 梁薇, 林直, 郭政宏. 2006. 兰花香味相关基因的RAPD分子标记. 江苏农业科学,(5):78-79.
[67]	Wang Mugui, Zeng Ruizhen, Xie Li, Gao Xuhua, Zhang Zhisheng. 2011. In vitro polyploid induction and its identification in Cymbidium sinense. Chinese Agricultural Science Bulletin, 27 (2):132-136. (in Chinese)
	王木桂, 曾瑞珍, 谢利, 高许花, 张志胜. 2011. 墨兰多倍体的离体诱导和鉴定. 中国农学通报, 27 (2):132-136.
[68]	Wang Wei. 2023. Origin investigation and definition discussion of‘Chinese Cymbidium’. Guangdong Landscape Architecture, 45 (5):18-21. (in Chinese)
	王巍. 2023. ‘中国兰花’词汇溯源及定义探讨. 广东园林, 45 (5):18-21.
[69]	Wei Yonglu, Jin Jianpeng, Gao Jie, Zheng Muxian, Lu Chuqiao, Yang Fengxi, Zhu Genfa. 2019. A new Cymbidium sinense cultivar‘Yinchunsu’. Acta Horticulturae Sinica, 46 (S2):2874-2875. (in Chinese)
	魏永路, 金建鹏, 高洁, 郑睦銛, 陆楚桥, 杨凤玺, 朱根发. 2019. 墨兰新品种‘迎春素’. 园艺学报, 46 (S2):2874-2875.
[70]	Wu Yingxiang. 1993. Chinese Cymbidium. 2nd ed. Beijing: China Forestry Publishing House. (in Chinese)
	吴应祥. 1993. 中国兰花. 2版. 北京: 中国林业出版社
[71]	Wu Yingxiang. 1995. What is‘Chinese Cymbidium’and‘Exotic orchid’? Flower and bonsai(Flower Horticulture),(4):6. (in Chinese)
	吴应祥. 1995. 什么是“国兰”?什么是“洋兰”? 花木盆景(花卉园艺),(4):6.
[8]	Chen Xinqi. 2008. Delicate scent orchid in a hollow valley. Garden,(12):120-121. (in Chinese)
	陈心启. 2008. 空谷幽兰. 园林,(12):120-121.
[9]	Chen Xinqi. 2011. Manual of Chinese Cymbidium and their cultivars. Beijing: China Forestry Publishing House. (in Chinese)
	陈心启. 2011. 国兰及其品种全书. 北京: 中国林业出版社.
[10]	Chen Rumeng, Peng Mengxuan, Wei Qian, Zeng Ruizhen, Su Qinglian, Zhang Zhisheng. 2022. Factors influencing on the induction and development of testtube flower in Cymbidium. Molecular Plant Breeding, https://kns.cnki.net/kcms/detail/46.1068.S.20220829.1026.002.html. in Chinese)
	陈如梦, 彭梦萱, 魏倩, 曾瑞珍, 宿庆连, 张志胜. 2022. 影响兰花试管花诱导和发育的因素. 分子植物育种, https://kns.cnki.net/kcms/detail/46.1068.S.20220829.1026.002.html.
[11]	Cheng Fenfang, Chen Xinrong, Chen Yingying, Chen Gang. 2015. Research progress of tissue culture in Cymbidium sinense. Heilongjiang Agricultural Sciences,(3):155-159. (in Chinese)
	程芬芳, 陈新荣, 陈莹莹, 陈刚. 2015. 墨兰的组织培养研究进展. 黑龙江农业科学,(3):155-159.
[12]	Chu Yunxia, Zhang Yongchun, Jing Xiangmi. 2009. The method for radiation and EMS induced mutationon Cymbidium goeringii rhizomes in vitro. Northern Horticulture,(1):49-52. (in Chinese)
	褚云霞, 张永春, 靖相密. 2009. 春兰离体根状茎辐射及EMS诱变方法建立. 北方园艺,(1):49-52.
[13]	Feng Guomei, Li Heng. 1980. A new species of Orchidaceae from Cymbidium. Acta Botanica Yunnanica, 2 (3):334-336. (in Chinese)
	冯国楣, 李恒. 1980. 兰属的一个新种. 云南植物学报, 2 (3):334-336.
[14]	Fu S B, Zhou Z, Yang Y P, Ying Z, Xu W. 2023. Hormonal profiling of encapsulated and nonencapsulated rhizomes of Chinese Cymbidium in different storage. Environments Journal of Plant Growth Regulation,42:1006-1016.
[15]	Fu Xuelin, Zhang Zhisheng, He Ping, Ou Xiujuan, He Qiongying. 2000. The effect of ⁶⁰Coγ-ray irradiation on the growth and regeneration of Cymbidium sinense rhizome. Acta Agriculturae Nucleatae Sinica, 14 (6):333-336. (in Chinese)
	傅雪琳, 张志胜, 何平, 欧秀娟, 何琼英. 2000. ⁶⁰Coγ射线辐照对墨兰根状茎生长和分化的效应研究. 核农学报, 14 (6):333-336.
[16]	Gao R, Wu S Q, Piao X C, Park S Y, Lian M L. 2014. Micropropagation of Cymbidium sinense using continuous and temporary airlift bioreactor systems. Acta Physiologiae Plantarum, 36 (1):117-124.
[72]	Wu Yingxiang, Chen Xinqin. 1966. Three species new species of Cymbidium from provincia Szechuan. Acta Phytotaxonomica Sinica, 11 (1):31-34. (in Chinese)
	吴应祥, 陈心启. 1966. 四川兰属植物三新种. 植物分类学报, 11 (1):31-34.
[73]	Wu Yingxiang, Chen Xinqin. 1991. Two new species of Cymbidium from China. Acta Phytotaxonomica Sinica, 29 (6):549-552. (in Chinese)
	吴应祥, 陈心启. 1991. 中国兰属植物新两种. 植物分类学报, 29 (6):549-552.
[74]	Wu Z J, Liu S J, An B J, Zhang H, Wu J J, Li C F, Long Y. 2024. Mutagenesis and flowering promotion through sodium azide in vitro culture of Cymbidium faberi Rolfe. Horticulturae,10:899.
[75]	Xie Li, Wang Fen, Zeng Ruizhen, Guo Herong, Zhou Yuliang, Zhang Zhisheng. 2015. Agrobacterium-mediated transformation of Cymbidium sinense. Chinese Journal of Biotechnology, 31 (4):542-551. (in Chinese)
	谢利, 王芬, 曾瑞珍, 郭和蓉, 周玉亮, 张志胜. 2015. 农杆菌介导的墨兰遗传转化. 生物工程学报, 31 (4):542-551.
[76]	Xu Dongsheng. 2000. Appreciation and cultivation of famous Cymbidium ensifolium varieties in China. Beijing: China Forestry Publishing House. (in Chinese)
	许东生. 2002. 中国建兰名品赏培. 北京: 中国林业出版社..
[77]	Xu Dongsheng. 2002. Appreciation and cultivation of famous Cymbidium sinense varieties in China. Beijing: China Forestry Publishing House. (in Chinese)
	许东生. 2002. 中国墨兰名品赏培. 北京: 中国林业出版社..
[78]	Xu Dongsheng. 2003. Appreciation and cultivation of famous Cymbidium kanran varieties in China. Beijing: China Forestry Publishing House. (in Chinese)
	许东生. 2003. 中国寒兰名品赏培. 北京: 中国林业出版社..
[79]	Xu Dongsheng. 2003. Appreciation and cultivation of famous Cymbidium tortisepalum varieties in China. Beijing: China Forestry Publishing House. (in Chinese)
	许东生. 2003. 中国莲瓣兰名品赏培. 北京: 中国林业出版社..
[80]	Xu Dongsheng. 2007. Appreciation and cultivation of famous Cymbidium goeringii varieties in China. Shenyang: Liaoning Science and Techinology Publishing House. (in Chinese)
	许东生. 2007. 中国春兰名品赏培. 沈阳: 辽宁科学技术出版社.
[17]	Geng Qingzhi. 2015. Establishment of technique system of mutation breeding and mutant creation in Cymbidium‘Junhao’[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	耿庆芝. 2015. 君豪兰诱变技术体系的建立和突变体筛选[硕士论文]. 广州: 华南农业大学.
[18]	Guo Shuwei. 2016. Lan Pu. Zhengzhou: Zhongzhou Ancient Books Publishing House. (in Chinese)
	郭树伟. 2016. 兰谱. 郑州: 中州古籍出版社.
[19]	He Yuanhao, Hu Fengrong, Sun Chongbo, Chen Yue, Wang Yunzhu. 2022. A new Chinese Cymbidium cultivar‘Huangyi’. Acta Horticulturae Sinica, 49 (S1):125-126. (in Chinese)
	何元浩, 胡凤荣, 孙崇波, 陈跃, 王筠竹. 2022. 国兰新品种‘黄逸’. 园艺学报, 49 (S1):125-126.
[20]	Huang W T, Fang Z M, Zeng S J, Zhang J X, Wu K L, Chen Z L,Teixeira da Silva J A,Duan J. 2012. Molecular cloning and functional analysis of three FLOWERING LOCUS T (FT) homologous genes from Chinese Cymbidium. International Journal of Molecular Sciences, 13 (9):11385-11398.
[21]	Huang W T, Huang X Y, Fang Z M. 2022. Melatonininduced rhizome proliferation,differentiation,and rooting during rapid propagation of Cymbidium goeringii and Cymbidium faberi. HortScience, 57 (9):1072-1077.
[22]	Huang Yan, Li Fang, Chai Mingliang, Xia Yiping, Chen Kunsong. 2009. Effects of ⁶⁰Coγ-rays on chromosome ploidy and related enzyme activities in rhizomes of Cymbidium goeringii. Acta Agriculturae Nucleatae Sinica, 23 (6):971-975. (in Chinese)
	黄焱, 李方, 柴明良, 夏宜平, 陈昆松. 2009. ⁶⁰Coγ射线对春兰根状茎染色体倍性及相关酶活性的影响. 核农学报, 23 (6):971-975.
[23]	Jia Sisi, Zeng Ruizhen, Zhang Zhisheng, Wei Qian, Xie Li, Guo Herong. 2024. Research advances on molecular breeding technique of orchid. Journal of South China Agricultural University, 45 (1):1-14. (in Chinese)
	贾思思, 曾瑞珍, 张志胜, 魏倩, 谢利, 郭和蓉. 2024. 兰花分子育种技术研究进展. 华南农业大学学报, 45 (1):1-14.
[24]	Jiao Jinchuan, Zhu Gang. 2018. Research progress on tissue culture and variety breeding of Chinese Cymbidium. Southern Horticulture, 29 (4):61-64. (in Chinese)
	焦晋川, 朱刚. 2018. 国兰组织培养与育种研究进展. 南方园艺, 29 (4):61-64.
[25]	Jiang Bin, Lai Zhanglong. 2015. The breeding technique for the new variety Cymbidium goeringii‘Huayeshuangyi’. Fujian Forestry,(1):42-44. (in Chinese)
	江斌, 赖张龙. 2015. 兰花新品种“花叶双艺”的选育研究. 福建林业,(1):42-44.
[81]	Xu Dongsheng. 2009. Appreciation and cultivation of famous Cymbidium faberi varieties in China. Shenyang: Liaoning Science and Techinology Publishing House. (in Chinese)
	许东生. 2009. 中国蕙兰名品赏培. 沈阳: 辽宁科学技术出版社.
[82]	Xu Dongsheng. 2010. Appreciation and cultivation of famous Cymbidium tortisepalum var. longibracteatum varieties in China. Beijing: China Forestry Publishing House. (in Chinese)
	许东生 2010. 中国春剑兰名品赏培. 北京: 中国林业出版社.
[83]	Xu Lipu. 2018. Study on micropropagation technology of Cymbidium Jade Hare[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	徐礼谱. 2018. 玉兔兰组培快繁技术研究[硕士论文]. 广州: 华南农业大学.
[84]	Xu Wan, Lin Yajun, Zhao Zhuang, Zhou Zhuang. 2022. Advances in genetic resources and breeding research of Cymbidium. Acta Horticulturae Sinica, 49 (12):2722-2742. (in Chinese)
	徐婉, 林雅君, 赵荘, 周庄. 2022. 兰属植物资源与育种研究进展. 园艺学报, 49 (12):2722-2742.
[85]	Xu X Y, Ding C C, Hu W Q, Yu X, Zheng Y, Zhang D Y, Liao X Y, Liu X D, Liu Z J, Lan S R. 2021. Cymbidium xichouense(Orchidaceae;Epidendroideae),a new species from China:Evidence from morphological and molecular data. Phytotaxa, 484 (3):291-297.
[86]	Yang Andao. 1412. Nan Zhong You Fang Lu(Manuscript). (in Chinese)
	杨安道. 1412. 南中幽芳录(手抄本).
[87]	Yang F X, Gao J, Li J, Wei Y L, Xie Q, Jin J P, Lu C Q, Zhu W, Wong S K, Zhu G F. 2024. The China orchid industry:Past and future perspectives. Ornamental Plant Research,4:e002.
[88]	Yang Jing, Xie Li, Zeng Ruizhen, Shang Ruixin, Guo Herong, Zhang Zhisheng. 2016. Research on the factors affecting the proliferation,differentiation and seedling strength of rhizomes of Cymbidium‘Xiaofeng’. Northern Horticulture,(2):105-109. (in Chinese)
[26]	Jiang Yu. 2019. Creation and study on mechanism of the leaf color variation of Chinese Cymbidium[Ph. D. Dissertation]. Ya’an: Sichuan Agricultural University. (in Chinese)
	蒋彧. 2019. 一个国兰叶艺新材料的创制及叶艺形成机理研究[博士论文]. 雅安: 四川农业大学.
[27]	Jiang Yu, Tu Xunliang, He Junrong. 2023. Analysis of differential expression genes in leaves of leaf color mutant of Chinese orchid. Acta Horticulturae Sinica, 50 (2):371-381. (in Chinese)
	蒋彧, 涂勋良, 何俊蓉. 2023. 国兰叶色突变体叶片差异表达基因分析. 园艺学报, 50 (2):371-381.
[28]	Jin Jianpeng, Gao Jie, Xie Qi, Wei Yonglu, Lu Chuqiao, Yang Fengxi, Zhu Genfa. 2022. A new varietiy of Cymbidium sinense‘Sijihua’. Acta Horticulturae Sinica, 49 (S1):127-128. (in Chinese)
	金建鹏, 高洁, 谢琦, 魏永路, 陆楚桥, 杨凤玺, 朱根发. 2022. 墨兰新品种‘四季花’. 园艺学报, 49 (S1):127-128.
[29]	Kim M S, Won J Y, Song C H, Eun J S, Lee D W. 1997. Polyploidy induction of Cymbidium kanran by treatment of colchicine in vitro. RDA Journal of Horticulture Science, 39 (1):73-76.
[30]	Kong Fanlong. 2008. Tissue culture and establishing Agrobacterium-mediated transformation system in Cymbidium goeringii Rchb.f.[M. D. Dissertation]. Hangzhou: Zhejiang University. (in Chinese)
	孔凡龙. 2008. 春兰的组织培养与遗传转化的初步研究[硕士论文]. 杭州: 浙江大学.
[31]	Lee H Y, Jung J S, Lee J S. 1998. Induction of chlorophyll deficient mutant plant of Cymbidium kanran by EMS treatment. Korean Journal of Plant Tissue Culture, 25 (3):183-187.
[32]	Lee J, Kwon S, Park S, Kim D Y. 2015a. Development of new variety with a single fragrance flower in a peduncle and marginal stripe patterns in Cymbidium goeringii‘Ariul’. Korean Journal of Breeding Science, 47 (4):454-458.
[33]	Lee J, Kwon S, Park S, Kim D Y. 2015b. Development of new variety with multiple fragrance flowers in a peduncle and marginal stripe patterns in Cymbidium goeringii‘Julie’. Korean Journal of Breeding Science, 47 (4):459-463.
[34]	Lei Lei. 2017. Identification of gene related to the formation of 2n male gamete in Cymbidium and creation of polyploid germplasm resources[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	雷蕾. 2017. 兰花2n雄配子发生相关基因鉴定及多倍体资源创建[硕士论文]. 广州: 华南农业大学.
[88]	杨靖, 谢利, 曾瑞珍, 商瑞昕, 郭和蓉, 张志胜. 2016. 影响‘小凤兰’根状茎增殖、分化和壮苗的因素研究. 北方园艺,(2):105-109.
[89]	Yu Lu. 2022. Study on tissue culture and SRAP genetic diversity of Cymbidium goeringii‘HuRui’[M. D. Dissertation]. Hangzhou: Zhejiang University. (in Chinese)
	余璐. 2022. 春兰‘虎蕊’组织培养及SRAP遗传多样性研究[硕士论文]. 杭州: 浙江大学.
[90]	Zeng Ruizhen, Li Yanghui, Guo Herong, Xie Li, Su Qinglian, Zhu Jiao, Zhang Zhisheng. 2016. A new Cymbidium cultivar‘Xiaofeng’. Acta Horticulturae Sinica, 43 (S2):2817-2818. (in Chinese)
	曾瑞珍, 黎扬辉, 郭和蓉, 谢利, 宿庆连, 朱娇, 张志胜. 2016. 兰花新品种‘小凤兰’. 园艺学报, 43 (S2):2817-2818.
[91]	Zeng Ruizhen, Li Yanghui, Xie Li, Guo Herong, Yi Maosheng, Su Qinglian, Zhang Zhisheng. 2017. A new Chinese Cymbidium cultivar‘Yuxiang’. Acta Horticulturae Sinica, 44 (S2):2723-2724. (in Chinese)
	曾瑞珍, 黎扬辉, 谢利, 郭和蓉, 易懋升, 宿庆连, 张志胜. 2017. 国兰新品种‘玉香兰’. 园艺学报, 44 (S2):2723-2724.
[92]	Zeng Songjun, Wu Kunlin, Zhang Jianxia, Chen Zhilin, Duan Jun. 2013. Advance on research of breeding and hybridization of Chinese Cymbidium. Chinese Wild Plant Resources, 32 (1):1-5. (in Chinese)
	曾宋君, 吴坤林, 张建霞, 陈之林, 段俊. 2013. 国兰新品种的选育和杂交育种研究进展. 中国野生资源, 32 (1):1-5.
[93]	Zeng Wenjing. 2019. Studies on tissue culture of several Chinese Cymbidiums[M. D. Dissertation]. Beijing: Beijing Forestry University. (in Chinese)
	曾文静. 2019. 几种国兰组织培养技术研究[硕士论文]. 北京: 北京林业大学.
[94]	Zhang Fengrong. 1990. Cymbidium collection of Taiwan. Beijing: Science Press. (in Chinese)
	张丰荣. 1990. 台湾兰蕙集锦. 北京: 科学出版社.
[95]	Zhang Tianxiang. 2012. Study on the technique system of tissue culture in Cymbidium kanran Makino[M. D. Dissertation]. Fuzhou: Fujian Agriculture and Forestry University. (in Chinese)
	张天翔. 2012. 寒兰组织培养技术体系研究[硕士论文]. 福州: 福建农林大学.
[96]	Zhang Zhisheng. Orchid breeding// Liu Qinglin, Jia Guixia. Orchid breeding // Liu Qinglin,Jia Guixia. Landscape plant genetics and breeding. Third edition. Beijing:China Forestry Publishing House:394-423. (in Chinese)
	张志胜. 2024. 兰花育种 // 刘青林,贾桂霞. 园林植物育种学. 第3版. 北京: 中国林业出版社:394-423.
[97]	Zheng Gaixiao, Li Xia, Xiong Dongjin, Yang Boyun. 2017. ISSR analysis of the mutagenic effect on Cymbidium kanran in vitro. Journal of Nanchang University(Natural Science), 41 (3):255-258. (in Chinese)
	郑改笑, 李夏, 熊冬金, 杨柏云. 2017. 寒兰离体化学诱变效果的ISSR分析. 南昌大学学报(理科版), 41 (3):255-258.
[98]	Zheng Qi, Sun Yefang, Zhao Hu, Xing Hai, Huang Yuefu, Lin Guomei. 2011. A molecular specific marker of Cymbidium goeringii plum petal variety and its obtaining method. China:CN201010502030.4. (in Chinese)
	郑琪, 孙叶芳, 赵虎, 邢海, 黄岳夫, 林国梅. 2011. 一种春兰梅瓣品种的分子特异标记及其获得方法. 中国:CN201010502030.4.
[99]	Zhou Huiming, Mo Zhilong, Chen Changming, Lin Huifeng. 2022. A new Cymbidium cultivar‘Shayangcuidie’. Acta Horticulturae Sinica, 49 (12):2767-2768. (in Chinese)
	周辉明, 莫智龙, 陈昌铭, 林辉锋. 2022. 兰花新品种‘沙阳翠蝶’. 园艺学报, 49 (12):2767-2768.
[100]	Zhou Z, Ying Z, Wu Z G, Yang Y P, Fu S B, Xu W, Yao L J, Zeng A P, Huang J, Lan S R, Wang X L, Liu Z J. 2021. Anthocyanin genes involved in the flower coloration mechanisms of Cymbidium kanran. Frontiers in Plant Science,12:737815.
[101]	Zhu Genfa. 2016. Status and trend of orchid industrialization in China. China Flower & Gardening News. (in Chinese)
	朱根发. 2016. 中国兰花产业化现状与趋势.中国花卉报.
[102]	Zhu Genfa, Xu Yechun, Yin Zhengbao. 2020. Chinese Cymbidium and culture. Chinese Flower Horticulture,(13):18-21. (in Chinese)
	朱根发, 徐晔春, 尹正保. 2020. 国兰与文化. 中国花卉园艺,(13):18-21.
[103]	Zhu Guobing, Yang Baiyun, Ao Aiyan. 2008. Tissue culture and in viro flowering of Cymbidium kanran Makino. Plant physiology Journal, 44 (3):513-514. (in Chinese)
	朱国兵, 杨柏云, 熬爱艳. 2008. 寒兰的组织培养和试管开花. 植物生理学通讯, 44 (3):513-514.
[104]	Zhu Jiao. 2014. The occurrence of 2n male gamete and creation of sexual polyploid germplasm resource in orchids[Ph. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	朱娇. 2014. 兰花2n雄配子发生及有性多倍体资源创建[博士论文]. 广州: 华南农业大学.
[35]	Li C R, Dong N, Zhao Y M, Wu S S, Liu Z J, Zhai J W. 2021. A review for the breeding of orchids:Current achievements and prospects. Horticultural Plant Journal, 7 (5):380-392.
[36]	Li Hanyue. 2022. Study on technology of creation of Cymbidium ensifolium with high efficiency of micropropagation[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	李晗玥. 2022. 易工厂化繁殖建兰育种技术研究[硕士论文]. 广州: 华南农业大学.
[37]	Li Jingyuan. 2011. Radiationinduced effects on the rhizomes of Cymbidium kanran Makino in vitro after ⁶⁰Co-gamma ray radiation[M. D. Dissertation]. Nanchang: Nanchang University. (in Chinese)
	李婧源. 2011. ⁶⁰Coγ射线对寒兰根状茎的辐射诱变效应研究[硕士论文]. 南昌: 南昌大学.
[38]	Li M M, Su Q L, Zu J R, Xie L, Wei Q, Guo H R, Chen J J, Zeng R Z, Zhang Z S. 2022. Triploid cultivars of Cymbidium act as a bridge in the formation of polyploid plants. Frontiers in Plant Science,13:1029915.
[39]	Li Xiaohong. 2020. Study on technique of molecular marker-assisted flower color selection and identification of flower colour and fragrance gene in Cymbidium[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	李晓红. 2020. 兰花花香花色基因鉴定和分子标记辅助花色选择技术研究[硕士论文 ]. 广州: 华南农业大学.
[40]	Li Xiujuan, Zhu Weiyin, Chen Xiaochou, Lai Zhanglong, Jiang Ruirong, Gao Xiaokun, Xu Jianqiu. 2018. A new varietiy of Cymbidium‘Yunvdanxin’. Acta Horticulturae Sinica, 45 (S2):2799-2800. (in Chinese)
	李秀娟, 朱尾银, 陈孝丑, 赖张龙, 江瑞荣, 高小坤, 徐建球. 2018. 兰花新品种‘玉女丹心’. 园艺学报, 45 (S2):2799-2800.
[41]	Lin Bing, Chen Shilin, Huang Minling, Zhong Huaiqin, Chen Yuanquan, Wu Jianshe, Ye Xiuxian. 2009. Effects of ⁶⁰Coγ-rays irradiation on growth and development of three varieties of Chinese Cymbidium. Acta Agriculturae Nucleatae Sinica, 23 (2):244-247. (in Chinese)
	林兵, 陈诗林, 黄敏玲, 钟淮钦, 陈源泉, 吴建设, 叶秀仙. 2009. ⁶⁰Coγ射线辐照对3种国兰生长的影响. 核农学报, 23 (2):244-247.
[42]	Lin Fen, Deng Guochu. 1997. Studies on the inductive mutation of spring orchid. Journal of Hunan Agricultural University, 23 (4):39-43. (in Chinese)
	林芬, 邓国础. 1997. 春兰人工诱变的研究. 湖南农业大学学报, 23 (4):39-43.
[43]	Liu Tianmeng. 2017. Agrobacterium-mediated genetic transformation of hybrid Cymbidium‘Junhong’and‘Xiaofeng’[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	刘天梦. 2017. 农杆菌介导的‘君红兰’和‘小凤兰’遗传转化[硕士论文]. 广州: 华南农业大学.
[44]	Liu Y, Zhang H L, Guo H R, Xie L, Zeng R Z, Zhang X Q, Zhang Z S. 2017. Transcriptomic and hormonal analyses reveal that YUC-mediated auxin biogenesis is involved in shoot regeneration from rhizome in Cymbidium. Frontiers in Plant Science,8:1072.
[45]	Liu Zhongjian, Chen Xinqi, Ru Zhengzhong. 2006. The genus Cymbidium in China. Beijng: Science Press. (in Chinese)
	刘仲健, 陈心启, 茹正忠. 2006. 中国兰属植物. 北京: 科学出版社.
[46]	Long Qiangyu, Zeng Yanhua, Bu Zhaoyang. 2021. Research advances of Chinese Cymbidium flowering in vitro. Southern Horticulture, 32 (3):83-87. (in Chinese)
	龙蔷宇, 曾艳华, 卜朝阳. 2021. 国兰试管开花研究进展. 南方园艺, 32 (3):83-87.
[47]	Lu Jiahui, Cui Yongyi. 2024. Tissue culture technology of Cymbidiun goeringii‘Songmei’. Molecular Plant Breeding, https://link.cnki.net/urlid/46.1068.S.20241113.1526.012. in Chinese)
	卢佳卉, 崔永一. 2024. 春兰‘宋梅’组织培养技术. 分子植物育种, https://link.cnki.net/urlid/46.1068.S.20241113.1526.012.
[48]	Lu Sicong. 2004. Advocate artificial orchid. Forest  Humankind,(5):36-39. (in Chinese)
	卢思聪. 2004. 提倡人工培育国兰新品种. 森林与人类,(5):36-39.
[49]	Luo Weiyu. 2024. Screening of mutant and antagonistic fungusagainst stem rot disease in hybrid Chinese Cymbidium[M. D. Dissertation]. Guangzhou: South China Agricultural University. (in Chinese)
	罗维宇. 2024. 杂交国兰茎腐病抗性突变体和生防菌筛选[硕士论文]. 广州: 华南农业大学.
[50]	Luo Weiyu, Su Qinglian, Zeng Ruizhen, Yuan Yun, Wei Qian, Zhang Zhisheng. 2023. Advances in mutagenesis breeding research of orchids. Chinese Journal of Tropical Crops, 44 (11):2135-2148. (in Chinese)
	罗维宇, 宿庆连, 曾瑞珍, 袁赟, 魏倩, 张志胜. 2023. 兰花诱变育种研究进展. 热带作物学报, 44 (11):2135-2148.
[51]	Mo Lei. 2007. Collection of ancient Cymbidium books. Beijing: China Forestry Publishing House. (in Chinese)
	莫磊. 2007. 兰花古籍撷萃. 北京: 中国林业出版社.
[52]	Ning H J, Ao S Y, Fan Y R, Fu J X, Xu C M. 2018. Correlation analysis between the karyotypes and phenotypic traits of Chinese Cymbidium cultivars. Horticulture,Environment and Biotechnology, 59 (1):93-103.
[53]	Niu Tian, Zhang Lin, Wang Houxin, Li Chengxiu, Yu Yongchang, Sun Fang, Wang Changxian. 2013. Research advances of Chinese Cymbidium breeding. Shandong Agricultural Sience, 45 (7):129-132. (in Chinese)
	牛田, 张林, 王厚新, 李承秀, 于永畅, 孙芳, 王长宪. 2013. 国兰育种研究进展. 山东农业科学, 45 (7):129-132
[54]	Ou Jince. 1992. Ling Hai Lan Yan. Lu Ziqing translated. Guangzhou: Guangdong People’s Publishing House. (in Chinese)
	区金策. 1992. 岭海兰言. 鲁子清译. 广州: 广东人民出版社

种类 Variety	命名时间 Nominated time	开始育种时间 Beginning time of the breeding	参考文献 Reference
春兰Cymbidium goeringii (Rchb. f.) Rchb. F.	1852	1110年前 Before 1110	吴应祥,1993
蕙兰Cymbidium faberi Rolfe	1896	1110年前 Before 1110	吴应祥,1993
建兰Cymbidium ensifolium (L.) Sw.	1799	1233年前 Before 1233	郭树伟,2016
墨兰Cymbidium sinense (Jackson ex Andr.) Willd	1805	1233年前 Before 1233	郭树伟,2016
寒兰Cymbidium kanran Mokino	1902	1247年前 Before 1247	郭树伟,2016
莲瓣兰Cymbidium tortisepalum Fukuyama	1934	1412年前 Before 1412	杨安道,1412
峨眉春蕙Cymbidium omeiense Y. S. Wu & S. C. Chen	1966	1966年后 After 1966	吴应祥和陈心启,1966
丘北东蕙兰Cymbidium qiubeiense K. M. Feng et H. Li	1980	1980年后 After 1980	冯国楣和李恒,1980
珍珠矮Cymbidium nanulum Y. S. Wu et S. C. Chen	1991	1991年后 After 1991	吴应祥和陈心启,1991
西畴兰Cymbidium xichouense X. Y. Xu，C. C. Ding et S. R. Lan	2021	2021年后 After 2021	Xu et al.,2021

种类 Variety	命名时间 Nominated time	开始育种时间 Beginning time of the breeding	参考文献 Reference
春兰Cymbidium goeringii (Rchb. f.) Rchb. F.	1852	1110年前 Before 1110	吴应祥,1993
蕙兰Cymbidium faberi Rolfe	1896	1110年前 Before 1110	吴应祥,1993
建兰Cymbidium ensifolium (L.) Sw.	1799	1233年前 Before 1233	郭树伟,2016
墨兰Cymbidium sinense (Jackson ex Andr.) Willd	1805	1233年前 Before 1233	郭树伟,2016
寒兰Cymbidium kanran Mokino	1902	1247年前 Before 1247	郭树伟,2016
莲瓣兰Cymbidium tortisepalum Fukuyama	1934	1412年前 Before 1412	杨安道,1412
峨眉春蕙Cymbidium omeiense Y. S. Wu & S. C. Chen	1966	1966年后 After 1966	吴应祥和陈心启,1966
丘北东蕙兰Cymbidium qiubeiense K. M. Feng et H. Li	1980	1980年后 After 1980	冯国楣和李恒,1980
珍珠矮Cymbidium nanulum Y. S. Wu et S. C. Chen	1991	1991年后 After 1991	吴应祥和陈心启,1991
西畴兰Cymbidium xichouense X. Y. Xu，C. C. Ding et S. R. Lan	2021	2021年后 After 2021	Xu et al.,2021

品种名 Name of cultivar	父母本 Parents	参考文献/来源Reference/ Source
明若轻溪 Cym. georingii‘Mingruo Qingxi’	春兰‘黄荷梅’× 春兰‘黄牡丹奇’ Cym. georingii‘Huanghemei’× Cym. georingii‘Huangmu Danqi’	MARA
凤岗春树 Cym. georingii‘Fenggang Chunshu’	春兰‘黄荷梅’× 春兰‘牡丹奇’ Cym. georingii‘Huanghemei’× Cym. georingii‘Mudanqi’	MARA
盛世蝴蝶 Cym. georingii‘Shengshi Hudie’	春兰‘大富贵’× 春兰‘余蝴蝶’ Cym. georingii‘Dafugui’× Cym. georingii‘Yuhudie’	MARA
扬红梅 Cym.‘Yanghongmei’	春兰‘大富贵’× 春兰‘瑞梅’ Cym. georingii‘Dafugui’× Cym. georingii‘Ruimei’	孙叶等,2018
黄蝴蝶 Cym. goeringii‘Yellow butterfly’	春兰‘黄水仙’× 春兰‘余蝴蝶’ Cym. georingii‘Huangshuixian’× Cym. georingii‘Yuhudie’	包建忠等,2017
花叶双艺 Cym.‘Huaye Shuangyi’	春兰‘大富贵’× 春剑‘边草’ Cym. georingii‘Dafugui’× Cym. tortisepalum var. longibracteatum‘Biancao’	江斌和赖张龙,2015
点绛唇Cym.‘Dianjiangchun’	春兰‘大富贵’×‘素君荷’Cym. georingii‘Dafugui’× Cym.‘Sujunhe’	MARA
金富贵Cym.‘Jinfugui’	春兰‘簪蝶’×‘大富贵’Cym. georingii‘Zandie’× Cym.‘Dafugui’	ARDZ
福娃梅Cym.‘Fuwamei’	春兰‘宋梅’×‘大富贵’Cym. georingii‘Songmei’× Cym.‘Dafugui’	孙崇波等,2012
沙阳玲珑Cym.‘Shayang Linglong’	蕙兰‘大一品’×‘寒香梅’Cym. faberi‘Dayipin’× Cym.‘Hanxiangmei’	MARA
沙阳碧玉 Cym.‘Shayang Biyu’	蕙兰‘金奥素’× 墨兰‘企剑白墨’ Cym. faberi‘Jinaosu’× Cym. sinense‘Qijian Baimo’	MARA
富贵花开Cym.‘Fugui Huakai’	蕙兰‘金奥素’×‘素君荷’Cym. faberi‘Jinaosu’× Cym.‘Sujunhe’	MARA
金秋荷Cym. ensifolium‘Jinqiuhe’	建兰‘君荷’× 建兰‘黄荷’Cym. ensifolium‘Junhe’× Cym. ensifolium‘Huanghe’	MARA
黄逸Cym. ensifolium‘Huangyi’	建兰‘君荷’× 建兰‘黄荷’Cym. ensifolium‘Junhe’× Cym. ensifolium‘Huanghe’	何元浩等,2022
粉蝶Cym. ensifolium‘Fendie’	建兰‘市长红’× 建兰‘大宝岛’Cym. ensifolium‘Shizhanghong’× Cym. ensifolium‘Dabaodao’	MARA
森彩奇洛Cym. ensifolium‘Sencai Qiluo’	建兰‘市长红’× 建兰‘富山奇蝶’Cym. ensifolium‘Shizhanghong’× Cym. ensifolium‘Fushan Qidie’	MARA
弄彩Cym. ensifolium‘Nongcai’	建兰‘四季双艺’× 建兰‘连城红’Cym. ensifolium‘Siji Shuangyi’× Cym. ensifolium‘Lianchenghong’	MARA
毓公主Cym. ensifoliumYugongzhu’	建兰‘荷王’× 建兰‘一品梅’Cym. ensifolium‘Hewang’× Cym. ensifolium‘Yipingmei’	GXZF
馨公主 Cym. ensifolium‘Xingongzhu’	建兰‘小国魂’× 建兰‘黄一品’ Cym. ensifolium‘Xiaoguohun’× Cym. ensifolium‘Huangyipin’	GXZF
仙野佛光Cym.‘Xianye Foguang’	建兰‘黄荷’×‘素君荷’Cym. ensifolium‘Huanghe’× Cym. ‘Sujunhe’	MARA
素玉Cym.‘Suyu’	建兰‘银边玉女素’×‘素君荷’Cym. ensifolium‘Yinbianyunüsu’× Cym.‘Sujunhe’	MARA
蝶舞Cym.‘Diewu’	建兰‘宝岛仙女’×‘素君荷’Cym. ensifolium‘Baodao Xiannü’× Cym.‘Sujunhe’	MARA
将相和Cym.‘Jiangxianghe’	建兰‘一品梅’×‘素君荷’Cym. ensifolium‘Yipinmei’× Cym.‘Sujunhe’	MARA
韵玉Cym.‘Yunyu’	建兰‘连城红’× 莲瓣兰‘小雪素’Cym. ensifolium‘Lianchenghong’× Cym. tortisepalum‘Xiaoxuesu’	MARA
玉女丹心Cym.‘Yunü Danxin’	建兰‘冠年’× 莲瓣兰‘白雪公主’Cym. ensifolium‘Guannian’× Cym. tortisepalum‘Baixue Gongzhu’	李秀娟等,2018
沙阳奇蝶Cym.‘Shayang Qidie’	墨兰‘富贵’× 春兰‘余蝴蝶’Cym. sinense‘Fugui’× Cym. georingii‘Yuhudie’	MARA
沙阳翠蝶 Cym.‘Shayangq Cuidie’	墨兰‘富贵’× 春兰‘余蝴蝶’ Cym. sinense‘Fugui’× Cym. georingii‘Yuhudie’	周辉明等,2022
紫韵玉兔兰Cym.‘Ziyun Yutu’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
粉墨玉兔兰Cym.‘Fenmo Yutu’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉坤兰Cym.‘Yukun’	墨兰‘金嘴’× 兔耳兰‘文引1号’ Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉锦兰Cym.‘Yujin’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	ARDG
玉姬兰 Cym.‘Yuji’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉澄兰Cym.‘Yucheng’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
桃花仙子 Cym.‘Taohua Xianzi’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	ARDG
玉香兰Cym.‘Yuxiang’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	曾瑞珍等,2017
玉叶兰Cym.‘Yuye’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉影兰Cym.‘Yuying’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉亭兰Cym.‘Yuting’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉鑫兰Cym.‘Yuxin’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin No.1’	MARA
君玉兰Cym.‘Junyu’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉缘兰Cym.‘Yuyuan’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶翠晶兰Cym.‘Moye Cuijing’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶紫晶兰Cym.‘Moye Zijing’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶紫玉兰Cym.‘Moye Ziyu’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	ARDG
小凤兰Cym.‘Xiaofeng’	墨兰‘企剑白墨’× 大凤兰Cym. sinense‘Qijian Baimo’× Cym.‘Dafeng’	曾瑞珍等,2016
国凤兰Cym.‘Guofeng’	墨兰‘企剑白墨’×‘小凤兰’Cym. sinense‘Qijian Baimo’× Cym.‘Xiaofeng’	ARDG
君豪兰Cym.‘Junhao’	墨兰‘企剑白墨’× 大凤兰Cym. sinense‘Qijian Baimo’× Cym.‘Dafeng’	MARA
凝香公主Cym.‘Ningxiang Gongzhu’	墨兰‘企剑黑墨’× 君豪兰Cym. sinense‘Qijian Heimo’× Cym.‘Junhao’	MARA
紫盈夫人Cym.‘Ziying Furen’	墨兰‘企剑黑墨’× 君豪兰Cym. sinense‘Qijian Heimo’× Cym.‘Junhao’	MARA
吕峰晴雪Cym.‘Lüfeng Qingxue’	寒兰‘寒素三星’×‘素君荷’Cym. kanran‘Hansu Sanxing’×Cym.‘Sujunhe’	MARA
馥公主Cym.‘Fugongzhu’	寒兰‘红香妃’× 建兰‘红宋梅’Cym. kanran‘Hongxiangfei’× Cym. ensifolium‘Hongsongmei’	GXZF
小王子Cym.‘Xiaowangzi’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	GXAAS
玉公主Cym.‘Yugongzhu’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	GXAAS
香公主Cym.‘Xianggongzhu’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	卜朝阳等,2019

品种名 Name of cultivar	父母本 Parents	参考文献/来源Reference/ Source
明若轻溪 Cym. georingii‘Mingruo Qingxi’	春兰‘黄荷梅’× 春兰‘黄牡丹奇’ Cym. georingii‘Huanghemei’× Cym. georingii‘Huangmu Danqi’	MARA
凤岗春树 Cym. georingii‘Fenggang Chunshu’	春兰‘黄荷梅’× 春兰‘牡丹奇’ Cym. georingii‘Huanghemei’× Cym. georingii‘Mudanqi’	MARA
盛世蝴蝶 Cym. georingii‘Shengshi Hudie’	春兰‘大富贵’× 春兰‘余蝴蝶’ Cym. georingii‘Dafugui’× Cym. georingii‘Yuhudie’	MARA
扬红梅 Cym.‘Yanghongmei’	春兰‘大富贵’× 春兰‘瑞梅’ Cym. georingii‘Dafugui’× Cym. georingii‘Ruimei’	孙叶等,2018
黄蝴蝶 Cym. goeringii‘Yellow butterfly’	春兰‘黄水仙’× 春兰‘余蝴蝶’ Cym. georingii‘Huangshuixian’× Cym. georingii‘Yuhudie’	包建忠等,2017
花叶双艺 Cym.‘Huaye Shuangyi’	春兰‘大富贵’× 春剑‘边草’ Cym. georingii‘Dafugui’× Cym. tortisepalum var. longibracteatum‘Biancao’	江斌和赖张龙,2015
点绛唇Cym.‘Dianjiangchun’	春兰‘大富贵’×‘素君荷’Cym. georingii‘Dafugui’× Cym.‘Sujunhe’	MARA
金富贵Cym.‘Jinfugui’	春兰‘簪蝶’×‘大富贵’Cym. georingii‘Zandie’× Cym.‘Dafugui’	ARDZ
福娃梅Cym.‘Fuwamei’	春兰‘宋梅’×‘大富贵’Cym. georingii‘Songmei’× Cym.‘Dafugui’	孙崇波等,2012
沙阳玲珑Cym.‘Shayang Linglong’	蕙兰‘大一品’×‘寒香梅’Cym. faberi‘Dayipin’× Cym.‘Hanxiangmei’	MARA
沙阳碧玉 Cym.‘Shayang Biyu’	蕙兰‘金奥素’× 墨兰‘企剑白墨’ Cym. faberi‘Jinaosu’× Cym. sinense‘Qijian Baimo’	MARA
富贵花开Cym.‘Fugui Huakai’	蕙兰‘金奥素’×‘素君荷’Cym. faberi‘Jinaosu’× Cym.‘Sujunhe’	MARA
金秋荷Cym. ensifolium‘Jinqiuhe’	建兰‘君荷’× 建兰‘黄荷’Cym. ensifolium‘Junhe’× Cym. ensifolium‘Huanghe’	MARA
黄逸Cym. ensifolium‘Huangyi’	建兰‘君荷’× 建兰‘黄荷’Cym. ensifolium‘Junhe’× Cym. ensifolium‘Huanghe’	何元浩等,2022
粉蝶Cym. ensifolium‘Fendie’	建兰‘市长红’× 建兰‘大宝岛’Cym. ensifolium‘Shizhanghong’× Cym. ensifolium‘Dabaodao’	MARA
森彩奇洛Cym. ensifolium‘Sencai Qiluo’	建兰‘市长红’× 建兰‘富山奇蝶’Cym. ensifolium‘Shizhanghong’× Cym. ensifolium‘Fushan Qidie’	MARA
弄彩Cym. ensifolium‘Nongcai’	建兰‘四季双艺’× 建兰‘连城红’Cym. ensifolium‘Siji Shuangyi’× Cym. ensifolium‘Lianchenghong’	MARA
毓公主Cym. ensifoliumYugongzhu’	建兰‘荷王’× 建兰‘一品梅’Cym. ensifolium‘Hewang’× Cym. ensifolium‘Yipingmei’	GXZF
馨公主 Cym. ensifolium‘Xingongzhu’	建兰‘小国魂’× 建兰‘黄一品’ Cym. ensifolium‘Xiaoguohun’× Cym. ensifolium‘Huangyipin’	GXZF
仙野佛光Cym.‘Xianye Foguang’	建兰‘黄荷’×‘素君荷’Cym. ensifolium‘Huanghe’× Cym. ‘Sujunhe’	MARA
素玉Cym.‘Suyu’	建兰‘银边玉女素’×‘素君荷’Cym. ensifolium‘Yinbianyunüsu’× Cym.‘Sujunhe’	MARA
蝶舞Cym.‘Diewu’	建兰‘宝岛仙女’×‘素君荷’Cym. ensifolium‘Baodao Xiannü’× Cym.‘Sujunhe’	MARA
将相和Cym.‘Jiangxianghe’	建兰‘一品梅’×‘素君荷’Cym. ensifolium‘Yipinmei’× Cym.‘Sujunhe’	MARA
韵玉Cym.‘Yunyu’	建兰‘连城红’× 莲瓣兰‘小雪素’Cym. ensifolium‘Lianchenghong’× Cym. tortisepalum‘Xiaoxuesu’	MARA
玉女丹心Cym.‘Yunü Danxin’	建兰‘冠年’× 莲瓣兰‘白雪公主’Cym. ensifolium‘Guannian’× Cym. tortisepalum‘Baixue Gongzhu’	李秀娟等,2018
沙阳奇蝶Cym.‘Shayang Qidie’	墨兰‘富贵’× 春兰‘余蝴蝶’Cym. sinense‘Fugui’× Cym. georingii‘Yuhudie’	MARA
沙阳翠蝶 Cym.‘Shayangq Cuidie’	墨兰‘富贵’× 春兰‘余蝴蝶’ Cym. sinense‘Fugui’× Cym. georingii‘Yuhudie’	周辉明等,2022
紫韵玉兔兰Cym.‘Ziyun Yutu’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
粉墨玉兔兰Cym.‘Fenmo Yutu’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉坤兰Cym.‘Yukun’	墨兰‘金嘴’× 兔耳兰‘文引1号’ Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉锦兰Cym.‘Yujin’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	ARDG
玉姬兰 Cym.‘Yuji’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
玉澄兰Cym.‘Yucheng’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	MARA
桃花仙子 Cym.‘Taohua Xianzi’	墨兰‘金嘴’× 兔耳兰‘文引1号’Cym. sinense‘Jinzui’× Cym. lancifolium‘Wenyin 1’	ARDG
玉香兰Cym.‘Yuxiang’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	曾瑞珍等,2017
玉叶兰Cym.‘Yuye’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉影兰Cym.‘Yuying’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉亭兰Cym.‘Yuting’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉鑫兰Cym.‘Yuxin’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin No.1’	MARA
君玉兰Cym.‘Junyu’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
玉缘兰Cym.‘Yuyuan’	墨兰‘小香’× 兔耳兰‘文引1号’Cym. sinense‘Xiaoxiang’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶翠晶兰Cym.‘Moye Cuijing’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶紫晶兰Cym.‘Moye Zijing’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	MARA
摩耶紫玉兰Cym.‘Moye Ziyu’	墨兰‘达摩’× 兔耳兰‘文引1号’Cym. sinense‘Damo’× Cym. lancifolium‘Wenyin 1’	ARDG
小凤兰Cym.‘Xiaofeng’	墨兰‘企剑白墨’× 大凤兰Cym. sinense‘Qijian Baimo’× Cym.‘Dafeng’	曾瑞珍等,2016
国凤兰Cym.‘Guofeng’	墨兰‘企剑白墨’×‘小凤兰’Cym. sinense‘Qijian Baimo’× Cym.‘Xiaofeng’	ARDG
君豪兰Cym.‘Junhao’	墨兰‘企剑白墨’× 大凤兰Cym. sinense‘Qijian Baimo’× Cym.‘Dafeng’	MARA
凝香公主Cym.‘Ningxiang Gongzhu’	墨兰‘企剑黑墨’× 君豪兰Cym. sinense‘Qijian Heimo’× Cym.‘Junhao’	MARA
紫盈夫人Cym.‘Ziying Furen’	墨兰‘企剑黑墨’× 君豪兰Cym. sinense‘Qijian Heimo’× Cym.‘Junhao’	MARA
吕峰晴雪Cym.‘Lüfeng Qingxue’	寒兰‘寒素三星’×‘素君荷’Cym. kanran‘Hansu Sanxing’×Cym.‘Sujunhe’	MARA
馥公主Cym.‘Fugongzhu’	寒兰‘红香妃’× 建兰‘红宋梅’Cym. kanran‘Hongxiangfei’× Cym. ensifolium‘Hongsongmei’	GXZF
小王子Cym.‘Xiaowangzi’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	GXAAS
玉公主Cym.‘Yugongzhu’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	GXAAS
香公主Cym.‘Xianggongzhu’	豆瓣兰 × 春兰Cym. serratum × Cym. goeringii	卜朝阳等,2019

方法 Method	材料 Material	结果 Result	参考文献 Reference
γ射线辐照 γ-ray irradiation	春兰幼苗 Seedlings of Cym. goeringii	5 Gy射线辐照的叶艺变异率为70.95%The rate of leaf color and pattern mutation irradiated by 5 Gy was 70.95%	孙叶等,2016
	春兰根状茎 Rhizome of Cym. goeringii	射线辐照适宜剂量为20 Gy，获得4个叶艺突变体The ⁶⁰Coγ-ray appropriate dose was 20 Gy，and four leaf color and pattern mutants were obtained	褚云霞等,2009
	蕙兰、建兰、墨兰成熟植株 Rhizome of Cym. Faberi，Cym. ensifolium and Cym. sinense	5 ~ 20 Gy 辐照处理中，筛选出2个兰新种质，其他变异体10个Two new germplasm resources and ten variants were screened from the radiation treatment of 5-20 Gy	包建忠等,2010
	建兰、墨兰成熟植株 Mature plant of Cym. ensifolium and Cym. sinense	10 ~ 20 Gy处理产生了叶片短小、叶艺、并蒂花、多花等突变类型Variants with short and small leaves，leaf color and pattern mutation，parallel or multiple flowers were produced in the irradiation treatment with 10-20 Gy	陈华等,2005
	建兰、墨兰根状茎 Rhizome of Cym. ensifolium and Cym. sinense	半致死量为10 ~ 20 Gy，得到叶艺株17株和矮化株5株The half-lethal radiation dose was 10-20 Gy，five dwarfs and seventeen mutants with leaf art and pattern were obtained	潘宏,2008
	建兰、墨兰成熟植株 Rhizome of Cym. ensifolium and Cym. sinense	半致死量为26 ~ 30 Gy，成熟假鳞茎耐辐照程度最大The half-lethal radiation dose was 26-30 Gy，the radiation tolerance of mature pseudobulbs was the highest	林兵等,2009
	墨兰根状茎 Rhizome of Cym. sinense	低剂量辐照促进根状茎增殖Low dose radiation promoted the proliferation of rhizome	傅雪琳等,2000
	寒兰根状茎 Rhizome of Cym. kanran	5 ~ 30 Gy辐射的根状茎生长良好，60 ~ 180 Gy辐射的根状茎生长基本停滞 The rhizomes irradiated with 5-30 Gy grew well，those irradiated with 60-180 Gy ceased to grow	李婧源,2011
	春剑幼芽 Buds of Cym. tortisepalum var. longibracteatum	20 Gy处理中获得了‘叶艺隆昌素’ Cym. tortisepalum var. longibracteatum‘Longchangsu’with leaf color and pattern mutation was obtained in the treatment of 20 Gy	蒋彧,2019
¹²C⁶⁺重离子辐照 ¹²C⁶⁺ heavy ion irradiation	‘君豪兰’根状茎 Rhizome of Cym.‘Junhao’	90 Gy辐照下再生植株表型变异率达100%，产生了株型直立、植株粗壮、矮小、颜色变深或变浅、叶片数增多等12个变异体 The phenotypic varation rate of irradiation with 90 Gy was 100%，twelve phenotypic variants including plantlets being erect，stout，malformed or tiny，with shallow or dark in colour，more and wider leaves etc.	耿庆芝,2015
¹²C⁶⁺重离子辐照 ¹²C⁶⁺ heavy ion irradiation	‘紫盈夫人兰’根状茎 Rhizome of Cym.‘Ziying Furen’	获得了1个抗茎腐病突变体 A mutation with resistance to stem rot disease was obtained	罗维宇,2024
紫外线辐照和秋水仙素处理 Ultraviolet ray irradiation and colchicine treatment	春兰原球茎 Protocorm of Cym. goeringii	产生矮化，叶片增宽增厚和叶艺变异体 Dwarf variant and variants with widened and thickened leaves and leaf color and pattern mutation were produced	林芬和邓国础,1997
EMS处理 EMS treatment	寒兰根状茎 Rhizome of Cym. kanran	0.2% EMS处理产生黄色线艺和白化植株等变异0.2% EMS treatment caused the variations including yellow linea in leaf and albino seedling	Lee et al.,1998
EMS处理 EMS treatment	春兰根状茎 Rhizome of Cym. goeringii	0.2% EMS处理产生矮化、白化、斑点和线艺叶艺突变体 Dwarfing，albino，mottling and linea mutants appeared in the regenerated plants with 0.2% EMS treatment	Shin et al.,2011
NaN₃处理 NaN₃ treatment	蕙兰类原球茎和根 Protocorm-like bodies (PLBs) and shoots of Cym. faberi	再生植株出现叶片较短、较厚、高度降低、叶片半透明等变异 Variations with shorter and thicker leaves，shorter node length，reduced height，and mid-translucent leaves appeared in the regenerants	Wu et al.,2024
化学诱变 Chemical mutagenesis	寒兰原球茎 Protocorm of Cym. kanran	七种化学试剂处理中DES最适合寒兰诱变 Among seven treatments of chemical mutagenesis reagent，DES is the most suitable for the mutation of Cym. kanran	郑改笑等,2017